Brachiopods of the Imtachan Formation in the Upper Permian of the Southern Verkhoyanie, Northeast Russia

R.V. Kutygin ^*, V.I. Makoshin ^**

Diamond and Precious Metal Geology Institute, Siberian Branch,
Russian Academy of Sciences, Yakutsk, 677000 Russia

E-mail: ^*rkutygin@mail.ru, ^**makoshin_89@mail.ru

Received December 22, 2022; Accepted May 26, 2023

ORIGINAL ARTICLE

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DOI: 10.26907/2542-064X.2023.2.308-321

For citation: Kutygin R.V., Makoshin V.I. Brachiopods of the Imtachan Formation in the Upper Permian of the Southern Verkhoyanie, Northeast Russia. Uchenye Zapiski Kazanskogo Universiteta. Seriya Estestvennye Nauki, 2023, vol. 165, no. 2, pp. 308–321. doi: 10.26907/2542-064X.2023.2.308-321. (In Russian)

Abstract

An extensive collection of fossil brachiopods, all assigned to the Wuchiapingian Stage, was recovered from the lower part of the Imtachan Formation along the Ustupnyi Creek (right tributary of the Setorym River) in the Southern Verkhoyanie. It comprises strophalosiids (Strophalosia? aff. vollossovitschi (Fredericks), Strophalosia? sp., Subtaeniothaerus lungersgauzeni Solomina, Marginalosia? magna Abramov et Grigorjeva), spiriferids (Crassispirifer monumentalis Abramov et Grigorjeva, Crassispirifer cf. monumentalis Abramov et Grigorjeva, Crassispirifer sp.), athyridids (Cleiothyridina nikolaevi Grunt, Cleiothyridina sp. 1, Bajtugania sp.), and terebratulids (Marinurnula? aff. mantuanensis (Campbell), Marinurnula? sp.). The identified assemblage is associated with brachiopods from the lower part of the Khalpirki Formation of the Western Verkhoyanie, the appearance of which marks the boundary between the Dulgalakhian and Khalpirkian Regional Stages and also substantiates the lower boundary of the Crassispirifer monumentalis Zone. Single finds of Crassispirifer cf. monumentalis in the lower part of the Middle Imtachan Subformation indicate that the entire lower subformation and the lower part of the middle subformation belong to the C. monumentalis Zone. There are no reliable finds of brachiopods at higher levels of the Imtachan Formation.

Keywords: Upper Permian, Wuchiapingian Stage, Khalpirkian Regional Stage, Imtachan Formation, biostratigraphy, brachiopods, Verkhoyanie

Acknowledgements. We are grateful to D. Bond, I.V. Brynko, S.S. Burnatny, A.S. Biakov, I.L. Vedernikov, V.I. Davydov, A.N. Kilyasov, and A.N. Naumov for participating in the joint field studies of the Imtachan Formation on the Ustupnyi Creek, as well as to M.F. Tretyakov and the staff of the Tompo Field Camp of NEFU led by him for their help in organizing the field work. The study was supported by the Russian Science Foundation and the Republic of Sakha (Yakutia) (project no. 22-24-20112, https://rscf.ru/en/project/22-24-20112/).

Figure Captions

Fig. 1. Section of the Imtachan Formation along the Ustupnyi Creek (on the left), its location and distribution of brachiopods (on the right): 1 – mudstones, 2 – fine-grained siltstones, 3 – coarse-grained siltstones, 4 – fine-grained sandstones, 5 – medium- and coarse-grained sandstones, 6 – diamictites, 7 – glendonites, 8 –concretions, 9 – brachiopods, 10 – ammonoids.

Fig. 2. Brachiopods of the lower part of the Imtachan Formation of the Ustupnyi Creek (collection of DPMGI): a–b – Marinurnula? aff. mantuanensis (Campbell), 202/511-13 and 202/511-15; c–d – Cleiothyridina nikolaevi Grunt, 202/516-53 and 202/516-63; e–f – Cleiothyridina sp. 1, 202/514-30 and 202/516-81; g – Strophalosia? aff. vollossovitschi (Fredericks), 202/513-1; h – Marginalosia? magna Abramov et Grigorjeva, 202/514-7; i–j – Subtaeniothaerus lungersgauzeni Solomina, 202/516-3 and 202/516-5; k–l – Crassispirifer monumentalis Abramov et Grigorjeva, 202/514-17 and 202/515-1; a–j, l – ventral valves, k – imprint of a ventral valve.

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