Chimarra manni Banks

Chimarra manni Banks

Figs. 20, 33, 105 –110

Chimarrha manni Banks, 1924: 448.

Chimarra indigota Mosely, 1941: 361, new synonym.

The genitalia of this species, particularly the gonopods, resemble those of C. helomyzida, C. kimminsi, C. cartwrighti, C. lavensis, C. devoensis, C. cakaudrovensis, C. devoensis, C. malaisei, and C. vanuensis. It is easily distinguished from C. kimminsi, C. helomyzida, C. tipulida and C. cartwrighti by the presence of 2 pairs of endothecal spines (not 1 pair); from C. lavensis, C. devoensis, and C. vanuensis by the absence of a prominent dorsad-oriented process on each of the lateral lobes of tergum X; and from C. cakaudrovensis by its shorter segment IX and more prominent dorsal branches of tergum X.

Male. Body (Fig. 33): Head and thorax dark brown, dorsal part of meso- and metathorax slightly darker brown than rest of thorax. Area between ocelli nearly black. Foreleg anterior claw as long as foreleg spur.

Wings (Fig. 20). Forewings 6.4 mm (lectotype), dark brown, iridescent. Forewings broad, ratio of length to breadth 3.0; R1 weakly undulating before crossvein r; radial sector not produced anterad immediately before discoidal cell; discoidal cell originating distally of mid-length of wing, 2.5x longer than wide; median cell slightly longer than discoidal cell; crossvein r originating from basis of R2; fork I originating immediately before crossvein s at distance equal to 1/5th length of crossvein s; nygma located near base of fork II; fork III 1 /6th as long as wing; fork V as long as fork II; Cu2 ending in wing margin well separated from A. Hind wings 5.3 mm (lectotype), brown, iridescent; ratio of length to breadth 2.8; margin weakly incurved at arculus, where Cu1 and Cu2 fused with margin; fork I originating from anterodistal corner of discoidal cell; fork III slightly longer than discoidal cell and 1/7th as long as wing; fork V longer than fork I; 1A+2A about 4x as long as 1A.

Male genitalia (Figs. 105–110). Segment IX nearly 1.2x taller than long; in lateral view each side produced anterad into rectangular ventral plate and 1/2 as high triangular dorsal plate; ventral margins slightly convex, slightly constricted at vertical apodeme; anterodorsal margin with deep, U-shaped concavity; each posterior margin slightly convex; segment IX ventrally with with setae restricted to band at posterior 1/3rd (Fig. 107). Dorsal part of segment IX irregular in lateral view, slightly membranous before tergum X. In dorsal view segment IX with long, narrow ventral plates; dorsal plates narrow, triangular; gap between dorsal plates oval. In ventral view segment IX narrowing strongly at mid-length at transverse apodeme, anterior margin shallowly triangular, posterior margin slightly concave. Tergum X short, divided into pair of lateral branches; in lateral view each lateral branch with broad dorsal, rounded lobe, and short, nearly straight, tube-shaped ventral branch, with 2 apical sensillae. Each ventral branch with ventrolaterad directed, sharply triangular lateral process situated at mid-length, near ventral margin. In dorsal and ventral view, ventral processes broad at basis; slender distally of lateral processes. Cerci located dorsally on posterior margin of segment IX, about at mid-height of tergum X; wart-like in lateral and dorsal view; with long setae. Gonopods nearly as long as height of segment IX; in lateral view broad basal 1/2, strongly expanded ventrally into rounded lobe, narrowing distally; distal 1/2 almost straight, nearly parallel-sided; each gonopod apparentlyunbranched in lateral view. In lateral view anterodorsal margin of each gonopod nearly straight, smooth; posteroventral margin undulating, with mesal megasetae on posterior margin of ventral lobe; apex with marginal, mesad oriented megasetae. In ventral view, gonopods slightly broader at base than at distal part, with nearly straight, slightly undulating posterior margins; gonopods slightly curved mesad toward rounded apex. Phallic apparatus longer than rest of genitalia (with expanded endotheca), straight along its length; narrowest at midlength in lateral view; in lateral view phallotheca about 2x thicker than narrowest posterior part; in ventral view about 2x thicker than posterior part; apicoventral spine absent; phallotremal sclerite with 4 posterior rays originating from anterior plate, central rays very thin, situated closely; 4 nearly black, variously directed endothecal spines present, all shorter than diameter of narrowest part of phallotheca; endothecal spicules absent.

Female. Unknown.

Lectotype male (here designated): “Fijis: Taviuni, Somo Somo. W. M. Mann ”; “ Type 14825”; “ Chimarrha manni Type Bks”. [no date]. Abdomen macerated, in glycerol in micro vial on same pin as rest of animal [MCZ].

Paralectotypes (here designated): same data as lectotype — 2 males.

Additional material examined: Type material of C. indigota (examined); TAVEUNI: Cakaudrove Prov., Mt. Koronibuabua, 3.2 km NW Lavena, Malaise trap, 4–18.i.2004, 16°51’16.9’’S, 179°53’26.2’’W [16.8547°S, 179.8906°W], 234 m, leg. E. Schlinger & M. Tokota’a [loc#41] — 11 males [NHRS]. Same data, except Malaise trap in rainforest, 24.x–19.xi.2003, 16°51’28.3’’S, 179°53’43.5’’W [16.8579°S, 179.8954°W], 233 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#42] — 3 males [FNIC]. Cakaudrove Prov., Lavena, Malaise trap, 4–19.xi.2003, 16°51.283’S, 179°53.370’W [16.8579°S, 179.8936°W], 212 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#43] — 1 male [BPBM]. Same data, except 16°51.322’S, 179°53.338’W [16.8553°S, 179.8890°W], 213 m [loc#40] — 1 male [BPBM]. Cakaudrove Prov., Soqulu House in Soqulu Estate, Malaise trap 1, 27.xii.2002 – 3.i.2003, 16.833°S, 180°E, 140 m, leg. E. Schlinger & M. Tokota’a [loc#44] — 2 males [FNIC]. VANUA LEVU: 0.3 km S Rokosalase Village, Malaise trap 1, 28.ix–15.xi.2004, 16.531°S, 179.019°E, 94 m, leg. I. Sakealevu [loc#30] — 2 males [BPBM]. Macuata Prov., Dogotuki, 2.5 km E Nasavu River, Malaise trap, 7.vii.2003, 16.2519°S, 179.7833°E, 226 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#28] — 1 male [BPBM]. Macuata Prov., Rokosalase, Malaise trap in forest, 8–21.vi.2004, 16°31.891’S, 179°01.147’E [16.5315°S, 179.0192°E], 105 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#27] — 1 male [BPBM]. Macuata Prov., 0.4 km S Rokosalase Village, Malaise trap in forest, 23.iv–8.v.2004, 16.532°S, 179.019°E, 118 m, leg. E. Schlinger & M. Tokota’a [loc#31] — 7 males [BPBM]. Macuata Prov., 0.6 km S Rokosalase Village, Malaise trap in forest, 23.iv–8.v.2004, 16.5333°S, 179.0181°E, 180 m, leg. E. Schlinger & M. Tokota’a [loc#26] 1 male [NHRS].

Distribution: Taveuni and Vanua Levu.