Eupogonius wappesi Lingafelter, Morris, Skillman, and Santos-Silva 2021, new species

Eupogonius wappesi Lingafelter, Morris, Skillman, and Santos-Silva, new species

(Fig. 26–29)

Description. Holotype male. Head capsule dark brown except reddish brown gulamentum; apex of palpomeres dark reddish brown; scape, pedicel, and antennomeres III–IV reddish brown basally, gradually reddish brown toward dark brown apex; antennomeres V–XI brown, slightly darkened apically. Pronotum dark brown except dark reddish brown areas with yellow pubescence (base, apex, and wide band on each side). Sides of prothorax dark brown. Prosternum dark reddish brown, except dark brown margins; prosternal process dark reddish brown in basal half, orangish in apical half. Mesoventral process mostly brown, with reddish brown central base and part of sides, dark brown close to procoxal cavities. Mesanepisternum, mesepimeron, and metanepisternum dark brown. Metaventrite dark brown, except dark reddish brown area near metacoxae. Elytra dark brown in dorsal third, except dark orangish brown area close to epipleural margin, gradually orangish brown toward apex. Procoxae dark brown; meso- and metacoxae partially orangish brown. Femora dark brown except reddish brown base. Tibiae orangish brown about basal third, dark brown toward apex. Tarsomeres dark orangish brown, with apex dark brown (especially tarsomere V). Abdominal ventrites dark reddish brown, with dark brown apex.

Head. Frons coarsely, densely punctate; with pale yellow pubescence close to eyes and clypeus, partially obscuring integument close to eyes, white, not obscuring integument on remaining surface; with long, erect yellowish setae interspersed centrally, and long, erect brownish setae close to eyes. Vertex and area behind upper eye lobes coarsely, moderately densely punctate; area between antennal tubercles and upper eye lobes, and area close to posterior margin of upper eye lobes with dense yellow pubescence; remaining surface of vertex and area behind upper eye lobes with very sparse, short yellowish white setae; with brownish setae interspersed (erect setae more abundant on yellow pubescent area of vertex). Area behind lower eye lobes coarsely, densely punctate; area close to eye with dense pale yellow pubescence (this area gradually widened toward ventral surface; remaining surface with short, sparse yellowish white setae; with a few long, erect brownish setae close to eye. Genae 0.4 times length of lower eye lobes; finely, sparsely punctate, except smooth apex; with yellowish white pubescence not obscuring integument, and a few long, erect brownish setae interspersed, except glabrous smooth area. Postclypeus with pale yellow pubescence partially obscuring integument, and long, erect brownish setae interspersed. Labrum coplanar with anteclypeus posteriorly, inclined anteriorly; with both, short and long dark setae directed forward posteriorly, and dense golden pubescence anteriorly. Gulamentum smooth, glabrous, depressed anteriorly. Distance between upper eye lobes 0.24 times distance between outer margins of eyes; in frontal view, distance between lower eye lobes 0.55 times distance between outer margins of eyes. Antennae 1.4 times elytral length, nearly reaching elytral apex. Scape coarsely, densely punctate; with whitish pubescence not obscuring integument, and long, erect, brown and yellowish white setae interspersed throughout (erect setae longer ventrally). Pedicel with whitish pubescence not obscuring integument, absent in dark area except at narrow apex; with long, erect brown setae interspersed throughout (erect setae longer ventrally). Antennomeres III–IV with whitish pubescence not obscuring integument in basal third, distinctly sparser, brownish on remaining surface; with long, erect, brown and yellowish white setae interspersed throughout (erect setae longer ventrally, and whitish setae more abundant dorsally). Antennomeres V–X with brownish pubescence distinctly not obscuring integument, short, erect white setae dorsally, and long, erect, both whitish and dark setae ventrally (dark setae more abundant). Antennomere XI with brownish pubescence not obscuring integument, and short, erect whitish setae interspersed (erect setae more abundant apically). Antennal formula based on antennomere III (only one male measured): scape = 0.85; pedicel = 0.25; IV = 1.08; V = 0.48; VI = 0.42; VII = 0.38; VIII = 0.38; IX = 0.32; X = 0.30; XI = 0.38.

Thorax. Prothorax wider than long; sides with conical tubercle about middle. Pronotum coarsely, abundantly punctate; with wide, longitudinal, dense yellow pubescent band on each side, from base to apex, transverse yellow pubescent band close to posterior margin, connecting lateral bands (narrowed centrally), and transverse yellow pubescent band near anterior margin, connecting lateral bands; remaining surface with white, sparse white pubescence; with long, erect brownish setae throughout. Sides of prothorax, coarsely, densely punctate; with dense yellow pubescence, except wide central area with sparse whitish pubescence; with long, erect brownish setae throughout. Prosternum coarsely, somewhat abundant punctate; with pale yellow pubescence partially obscuring integument, and long, erect setae of same color interspersed. Prosternal process with punctures slightly denser than on prosternum; basal half with pale yellow pubescence partially obscuring integument; posterior half with sparse yellowish pubescence, and long, erect yellowish setae apically; narrowest area 0.2 times procoxal cavity. Mesoventrite and mesoventral process with white pubescence not obscuring integument; mesanepisternum mostly with dense yellow pubescence; mesepimeron with yellowish pubescence, sparser than on mesanepisternum. Metanepisternum and sides of metaventrite with abundant yellowish pubescence not obscuring integument; remaining surface of metaventrite with abundant whitish pubescence not obscuring integument, except glabrous metathoracic discrimen; metaventrite coarsely, somewhat abundant punctate laterally (punctures finer, sparser toward metathoracic discrimen), and long, erect whitish setae interspersed. Scutellum with sparse yellowish white pubescence. Elytra. Coarsely, abundantly punctate in basal third, punctures gradually finer, sparser toward apex; with longitudinal yellow pubescent band dorsally, from base to apex, curved apically, another longitudinal yellow pubescent band close to epipleural margin, from base to apex, apically fused with dorsal yellow pubescent band; remaining surface with whitish pubescence not obscuring integument, slightly denser between two yellow pubescent bands; with long, erect, abundant brownish setae throughout. Legs. Femora with yellowish white pubescence not obscuring integument, except glabrous posterior 2/3 of ventral surface; with long, erect, yellowish white and brownish setae interspersed. Tibiae with abundant, bristly yellowish white pubescence partially obscuring integument, and long, erect both yellowish white and brownish setae interspersed. Metatarsomere I slightly shorter than II–III together.

Abdomen. Ventrite I with abundant yellowish white pubescence not obscuring integument, forming dense fringe apically; ventrites II–V with abundant yellowish white pubescence not obscuring integument, forming dense fringe apically on II–IV, except nearly glabrous area close to apical margin on II–IV, and long, decumbent, abundant yellow setae centrally on II–V; with long, erect brownish setae on sides of III–V, and posterior third of V; apex of ventrite V slightly rounded.

Dimensions (mm). Total length 5.45; prothoracic length 1.05; anterior prothoracic width 1.05; posterior prothoracic width 1.10; maximum prothoracic width 1.45; humeral width 1.70; elytral length 3.90.

Type material. Holotype male from MEXICO, QUINTANA ROO: Highway 307, 9 km S. Cancun, 08.VI.2005, F.W. Skillman Jr. (FSCA, formerly FWSC).

Etymology. The new species is dedicated to the late James E. Wappes, for his friendship and contributions to the knowledge of American Cerambycidae.

Remarks. Four species of Eupogonius LeConte, 1852 have a longitudinal elytral pubescent band dorsally from base to apex: E. cryptus Hovore, 1989, E. flavocinctus Bates, 1872, E. monzoni Wappes and Santos-Silva, 2020, and E. vittipennis Bates, 1885 (see photographs of these species in Bezark 2021). Eupogonius wappesi differs from E. cryptus and E. flavocinctus by the wider elytra, lateral tubercles of the prothorax distinctly more conspicuous, absence of longitudinal yellow pubescent band in center of vertex, and absence of yellow pubescent band in center of the pronotum (entirely covering vertex in E. flavocinctus); from E. monzoni by the wider elytra, absence of a longitudinal yellow pubescent band in the center of the pronotum, and absence of dense white pubescence on the elytra; and from E. vittipennis by the wider elytra, lateral tubercles of the prothorax distinctly more conspicuous, longitudinal yellow pubescent bands on the pronotum fused anteriorly and posteriorly (not fused in E. vittipennis), and absence of a longitudinal pubescent band on the elytral suture.