Cretapristocera longiscapa Jouault & Ngô-Muller & Pouillon & Nel 2021, SP. NOV.

CRETAPRISTOCERA LONGISCAPA SP. NOV.

(FIGS 2–4)

u r n: l s i d: z o o b a n k. o r g: a c t: 2 1 A 7 C A E C - 4 2 5 0 45A9-A3B3-28F099EAEC25.

Holotype: Female, accession number IGR.BU-009, nearly complete and well-preserved female specimen in a rectangular piece of amber measuring 9 × 7 × 2.5 mm, together with a small Hymenoptera: Chalcidoidea; housed in the amber collection of the Geological Department and Museum (IGR) of the University of Rennes, France.

Type locality and horizon: Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar; Lower Cenomanian, Upper Cretaceous.

Etymology: The specific epithet is derived from Latin longus, long, and scapus, shaft, for the elongated scape of this species.

Description: Female; body 2.1 mm long (as preserved), not depressed, almost smooth and not strongly foveolate; LFW 1.5 mm; castaneous; wings setose. LH 0.43 mm; WH 0.5 mm; WF 0.26 mm; HE 0.11 mm; OOL 0.25 mm; WOT 0.16 mm; DAO 0.04 mm; VOL 0.1 mm. Head not pubescent, with small, dorsal, median depression, finely punctuated but not sculptured; mandible with one strong and sharp tooth, projecting forward; six maxillary palpomeres, three (?) visible labial palpomeres; clypeus spined; antenna with ten flagellomeres with cylindrical crosssection; scape 0.37 × 0.07 mm, pedicel 0.13 × 0.04 mm (all measurements refer to length × width), F1 0.07 × 0.05 mm, F2 0.05 × 0.05 mm, F3 0.05 × 0.06 mm, F 4 0.06 × 0.07 mm, F5 0.06 × 0.08 mm, F 6 0.08 × 0.08 mm, F7 0.08 × 0.08 mm, F8 0.08 × 0.07 mm, F9 0.09 × 0.06 mm, F10 0.17 × 0.06 mm; occipital carina strongly marked; compound eye without microsetae; ocellar triangle not equilateral, with two posterior ocelli more distant from each other than from anterior. Mesosoma with dorsum smooth, not roughly sculptured; femora strongly thickened, tibia thin, tarsal claws simple; tibial spur formula 1-2-2; notauli well marked, posteriorly convergent; parapsidal furrows weak; metanotum developed medially and overlapping mesoscutellum posteriorly; metapectal–propodeal complex rectangular, not foveolate, with marginal carinae and no posterior spines; posterior margin of propodeum straight. Forewing covered with microsetae and bordered with small setae; anterior border not angularly incurved anterior to pterostigma; costal vein present; only veins C, Sc + R, Rs + M, 2r-rs&Rs, M + Cu, Cu and cells C (hardly visible), R and 2R1 present; vein cu-a absent and cell Cu opened; cell 2R1 distally opened; no basal stub of Rs + M; pterostigma long and narrow; poststigmal abscissa of R1 short, spectral veins not visible. Hindwing with only veins C and Sc + R and with three hamuli. Metasoma with short petiole; six tergites present (including three preserved detached from the body), T2 enlarged, convex, partly covering sternites; sting not preserved; length of clearly visible tergites: T1 0.23 mm, T2 0.29 mm, T3 at least 0.19 mm.

Remarks: It seems that the internal musculature of the fossil was preserved during the fossilization process, especially those of the legs (Figs 2–3). It could be interesting to compare the evolution of the musculature and the insertion of the muscles between the Cretaceous and extant Bethylidae in a future study, using 3D X-ray tomography.