Wanniyala orientalis Huber 2019, sp. n.

Wanniyala orientalis sp. n.

Figures 132–134, 165–171, 214

Wanniyala sp. (from Inginiyagala): Huber & Benjamin 2005: 3311, fig. 9.

Wanniyala SL38: Eberle et al. 2018 (molecular data). Huber et al. 2018: fig. 7.

Diagnosis. Males are distinguished from congeners by details of palp (retrolateral sclerite of procursus with short and wide dorsal process; distinctive processes of palpal trochanter, and distally strongly curved bulbal apophysis; Figs 165–168) and by modification of clypeus (pair of processes connected by protruding arc; similar only in W. mudita; cf. Figs 156–157). Females are easily distinguished from most known congeners (except W. hakgala, W. mudita) by shape of epigynum (strongly projecting and pointed in lateral view; Fig. 170); they differ from W. hakgala by large anterior angle between epigynal plate and abdomen (compare Fig. 170 and Huber & Benjamin 2005: fig. 5i); from W. hakgala and W. mudita also by details of internal genitalia (almost round pore plates; anterior valve narrower than in W. mudita; Figs 171, 214).

Etymology. The specific name is an adjective and refers to the distribution of this species in eastern Sri Lanka.

Type material. SRI LANKA: ♂ holotype, ZFMK (Ar 20078), North Central Province, Dimbulagala (7.860°N, 81.118°E), 140 m a.s.l., 11.iii.2017 (B.A. Huber).

Other material examined. SRI LANKA: 3♂ 5♀ 2 juvs, ZFMK (Ar 20079), and 4♀ in pure ethanol, ZFMK (SL124), same data as holotype. 1♂ 11♀ 1 juv., ZFMK (Ar 20080), and 3♀ 1 juv. in pure ethanol, ZFMK (SL122), North Central Province, Minneriya Forest (8.047°N, 80.832°E), 150 m a.s.l., 10.iii.2017 (B.A. Huber). 7♂ 2♀, ZFMK (Ar 20081), and 2♀ 1 juv. in pure ethanol, ZFMK (SL120), Central Province, Kandalama Forest (7.859°N, 80.711°E), 220 m a.s.l., 10.iii.2017 (B.A. Huber). 3♂ 9♀, ZFMK (Ar 20082), and 1♀ in pure ethanol, ZFMK (SL130), Uva Province, Inginiyagala (7.225°N, 81.535°E), 110 m a.s.l., 12.iii.2017 (B.A. Huber). 1♂, ZMUT (AA 3635), same locality, “in litter of stony djungle”, 21.xi.1972 (P. Lehtinen). 1♂ 1♀, NMSL, 4♂ 10♀, ZFMK (Ar 20083), and 3♀ 1 juv. in pure ethanol, ZFMK (SL139), Uva Province, near Okkampitiya (6.728°N, 81.336°E), 190 m a.s.l., 13.iii.2017 (B.A. Huber). 2♂ 5♀, ZFMK (Ar 20084), Uva Province, near Gowindahela (7.041°N, 81.538°E), 130–180 m a.s.l., 12.iii.2017 (B.A. Huber). 1♀, ZMUT (AA 3632), Moneragala District, “Gumbukana” [Kumbukana, ~ 6.805°N, 81.294°E], in teak litter, 19.xi.1972 (P. Lehtinen).

Description. Male (holotype). MEASUREMENTS. Total body length 1.9, carapace width 0.90. Distance PME- PME 140 µm, diameter PME 100 µm, distance PME-ALE 30 µm; AME absent. Sternum width/length: 0.66/0.54. Leg 1: 19.2 (4.6 + 0.4 + 4.8 + 7.5 + 1.9), tibia 2: 2.9, tibia 3: 2.2, tibia 4: 2.9; tibia 1 L/d: 69.

COLOR (in ethanol). Carapace ochre yellow with median and lateral dark bands; ocular area posteriorly dark; clypeus with pair of dark bands below eye triads; sternum black; legs ochre-yellow, with indistinct darker rings on femora (subdistally) and tibiae (proximally and subdistally); abdomen ochre-gray with dark marks dorsally and laterally, ventrally also with dark pattern.

BODY. Habitus as in Figs 132–133; ocular area slightly raised; carapace with shallow but distinct median furrow; clypeus with pair of processes connected by protruding arc (similar to W. mudita; cf. Figs 156–157); sternum unmodified.

CHELICERAE. Similar to W. mudita (cf. Figs 156–157), with pair of small apophyses proximally laterally and pair of long apophyses directed forwards, tips of apophyses distally directed downwards, distance between tips of apophyses: 0.76.

PALPS. In general very similar to known congeners (cf. Figs 150–155); coxa unmodified; trochanter with distinctive processes (Fig. 165); femur with indistinct small dorsal apophysis proximally; tibia very long (0.98); procursus with several distinctive elements (Figs 167–168); bulb with simple membranous embolus and strongly curved apophysis with rounded tip (Fig. 166).

LEGS. Without spines and curved hairs; with short vertical hairs in higher than usual density on all metatarsi (especially proximally); retrolateral trichobothrium on tibia 1 at 15%; prolateral trichobothrium absent on tibia 1; tarsus 1 with ~25 pseudosegments, distally distinct.

Male (variation). Tibia 1 in three other males from type locality: 4.3, 4.4, 4.6. Males from other localities have shorter legs: tibia 1 in 17 males from all other localities: 3.2–3.9 (mean 3.5). In males from Minneriya, the dorsal process of the retrolateral sclerite of the procursus (arrow in Fig. 168) is slightly smaller.

Female. In general similar to male (Fig. 134) but clypeus unmodified and legs with usual low number of short vertical hairs. Tibia 1 in 5 females from type locality: 3.7–4.0 (mean 3.9); in 38 females from all other localities: 2.6–3.7 (mean 3.1). Epigynum as in Figs 169–170, strongly protruding and pointed in lateral view; posterior plate apparently reduced(?) to two lateral sclerites mostly hidden behind epigynal plate and poorly visible in ventral view; internal genitalia as in Figs 171, 214, with narrow anterior ‘valve’ and pair of roundish pore plates.

Natural history. At Kandalama, the spiders were abundant under leaves and other objects on the forest floor; when disturbed, they did not vibrate but rather ran away. At Okkampitiya, the spiders were mainly found under loose bark on the ground.

Distribution. Widely distributed in eastern Sri Lanka, excluding high elevation areas (Fig. 226).