Cinnadenia Kosterm.

Genus Cinnadenia Kosterm.

Adansonia, sér. 2, 13: 223 (1973); Grierson & Long, Flora of Bhutan 1 (2): 39 (1984); Kochummen, Tree Flora of Malesia 4: 123 (1989); Rohwer, Families and Genera of Vascular Plants 2: 389 (1993); Van der Werff, Blumea 46: 135 (2001).

— Type species: Dodecadenia paniculata Hook.f. (1886) (synonym of Cinnadenia paniculata (Hook.f.) Kosterm.)

Litsea section Octolitsea, H.Liu, Bulletin de la Société botanique de France 80: 568 (1933).

— Type species: Litsea liyuyingii H.Liu (synonym of Cinnadenia liyuyingii (H.Liu) de Kok & Sengun, comb. nov.).

DISTRIBUTION. — Three species, one in North India and Bhutan, one in South China and North Vietnam and one in Peninsular Malaysia (see Fig. 1).

DESCRIPTION

Trees 3-30 m tall. Terminal leaf buds not perulate. Leaves alternate, venation pinnate, domatia absent. Inflorescence umbellate or a panicle or with 3 or 4-5 terminal flowers, axillary, rarely terminal, involucral bracts absent or present. Flowers unisexual, dimerous or trimerous; perianth lobes 4 or 6, equal, persistent, partly truncate or deciduous in fruit; tube short, funnel-shaped, enlarged and indurate in fruit. Male flowers, stamens 8-12, 18-26 or 32, 3 rd row with a pair of stalked or sessile glands on the filaments; anthers 4-celled, all introrse. Female flowers, ovary sessile, at the bottom of

perianth tube; style slender; stigma discoid or obscurely 3-lobed. Fruit a drupe, seated on a cup-shaped perianth tube (cupule), margin entire.

1. Leaves lanceolate; apex acuminate; upper surface smooth (hand lens), secondary venation hardly visible. Inflorescence with an apical whorl of 2-3 flowers; flowers with 4 perianth lobes.............. 2. C. malayana Kosterm.

— Leaves elliptic to lanceolate, sub-obovate or narrowly ovate; apex acute to indistinctly acuminate to obtuse; upper surface pitted (hand lens), secondary venation clearly visible. Inflorescence with apical whorl of 4-5 flowers or umbellate; flowers with 6 perianth lobes................................................................................................. 2

2. Twigs sparsely hairy to glabrous when young; terminal leaf bud 6.4-7.4 mm long, apex acuminate, glabrous to velutinous; flowers with 26 or 32 stamens.................. 1. C. liyuyingii (H.Liu) de Kok & Sengun, comb. nov.

— Twigs velutinous when young, glabrescent; terminal leaf bud 2.3-2.5 mm long, apex acute to indistinctly acuminate, velutinous; flowers with 9-12 stamens............................................ 3. C. paniculata (Hook. f.) Kosterm.