Alloxysta obscurata

Alloxysta obscurata (Hartig, 1840)

Figs 2.31, 5.3

Xystus obscuratus Hartig, 1840: 200. Type: deposited in ZSM (examined).

Allotria obscuratus – Taschenberg 1866: 130.

Allotria obscurata – Dalla Torre 1893: 34.

Dilyta obscurata – Kieffer 1900: 114.

Alloxysta (Alloxysta) obscurata – Dalla Torre & Kieffer 1902: 39.

Alloxysta obscurata – Andrews 1978: 87.

Diagnosis

Alloxysta obscurata is mainly characterized by a partially open radial cell that is 2.7 times as long as wide (Fig. 5.3); the presence of pronotal carinae; the absence of propodeal carinae; female antenna with

rhinaria beginning on F3, F1 longer than pedicel and F2, F2 subequal to F3, and F3 shorter than F4 (Fig. 2.31); male antenna with rhinaria beginning on F4, F2 slightly curved, F1 longer than pedicel and F2, F2 longer than F3, and F3 longer than F4. It is similar to A. macrophadna, but can be differentiated by the proportions of the flagellomere (F1 is longer than F2, F2 is shorter than F3, and F3 is shorter than F 4 in A. obscurata (Fig. 2.31), while F1 is subequal to F2, F2 is longer than F3, and F3 is subequal to F 4 in A. macrophadna (Fig. 2.25)), the shape of the flagellomere (no flagellomeres are curved in males of A. obscurata, but F2 and F3 are strongly curved in males of A. macrophadna), and the size of the radial cell (2.7 times as long as wide in A. obscurata (Fig. 5.3), but 3.0 times in A. macrophadna (Fig. 4.21)).

Material examined

Lectotype

GERMANY: ♀, [second from top, lectotype H.H. Evenhuis (orange label)], [Xystus obscuratus Hartig det. H.H. Evenhuis 1980], [Alloxysta obscurata (Hartig, 1840) ♀ M. Ferrer-Suay det. 2011] (ZSM).

Paralectotype

GERMANY: ♀, [Xystus obscuratus Hartig det. H.H. Evenhuis 1980], [first from top, Paralectotype Xystus obscuratus Hartig, 1840 ♀], [Alloxysta obscurata (Hartig, 1840) ♀ M. Ferrer-Suay det. 2011] (ZSM).

Additional specimens (9 ♂♂, 52 ♀♀)

AUSTRIA: 1 ♀ [Semmeringgebiet, Reichenan dist., 25 May–6 Jun. 1957, G.E.J. Nixon] (BMNH).

CROATIA: 1 ♀ [swept from vegetation by lake, Croatia, Plitvice, 4–10 Aug., Jugoslavia: 1955, R.L. Coe.] (BMNH B.M. 1955-460).

CZECH REPUBLIC: 3 ♀♀ [Moravia, Dyje River, near Znojmo, 12 Aug. 91, L. Masner] (CNCI C-263); 1 ♀ [Moravia, Lanzhot-Ranspurk, 7–9Aug. 1991, L. Masner, sweep, climax flood forest] (CNCI C-268); 2 ♀♀ [Moravia, 16 km N Blansko, 8 Aug. 1991, L. Masner, sweep. Tilia-Acer forest] (CNCI C-271); 1 ♂ [Moravia, Lanzhot-Ranspurk, 9–12 Aug. 1991, L. Masner, climax hardwood forest, PT] (CNCI C-270); 1 ♀ [Moravia, Ladnice environs 7–9 Aug. 1991, L. Masner riparian, forest] (CNCI C-272).

GERMANY: 3 ♀♀ [Schwarzwald, Todtmoos environs, 6 Aug. 1984, L. Masner, screen sweeping] (CNCI C-235).

JAPAN: 1 ♂ [Fukuoka, Mt. Hiko, 700 m, 9–10 May 1989, sweep, M.J. Sharkey] (CNCI C-183); 2 ♀♀ [Hokkaido, Sapporo, Jozankei, 350 m, 20–31 Jul. 1989, K. Maeto and M. Sharkey] (CNCI C-140); 4 ♀♀ [Hokkaido, Sapporo, Jozankei, 350 m, 20–31 Jul. 1989, K. Maeto and M. Sharkey] (CNCI C-151); 2 ♀♀ [Hokkaido, Sapporo, Jozankei, 350 m, Aug. 1989, K. Maeto and M. Sharkey, MT] (CNCI C-203); 1 ♂, 1 ♀ [Hokkaido, Furano Exp. Forest, 43°15′ N, 142°20′ E, 9 Aug. 1996, 500 m, L. Masner, sweep] (CNCI C-162); 3 ♀♀ [Hokkaido, Shibetsu-Rubesu, E. slope of Mt. Shari, 43°45′ N, 144°45′ E, 17 Aug. 1996, 700 m, L. Masner, sweep] (CNCI C-164); 1 ♀ [Hokkaido, Sapporo, Jozankei, 350 m, 20–31 Jul. 1989, MT, K. Maeto and M. Sharkey] (CNCI C-204); 3 ♀♀ [Hokkaido, Sapporo, Jozankei, 350 m, 10– 21 Aug. 1989, K. Maeto and M. Sharkey, MT] (CNCI C-206); 1 ♀ [Hokkaido, Nukabira, 600 m, 5 Jul. 1989, sweep, M.J. Sharkey] (CNCI C-215); 1 ♀ [Hokkaido, Sapporo, Jozankei, 350 m, 29 Aug.–12 Sep. 1989, K. Maeto and M. Sharkey, MT] (CNCI C-207).

POLAND: 1 ♀ [Przemysl, 12 Sep. 1984, N.D. Springate: 1 ♀] (BMNH).

RUSSIA: 1 ♂, 1 ♀ [Russian Far East, Primorskii Krai Lazovski Zapovednik, c. 170 Km E., Vladivostok, Lazo, 43°30′33″ N, 134°06′59″ E, 1375 m, 2 Jun.–3 Jul. 2001, Mountain top, malaise trap 458, M. Quest

coll., Alloxysta sp1., det. M. Forshage 2012] (BMNH B.M. 2009-59); 1 ♀ [Magadanskaya Oblast, Aborigen Field Station, 500 m, 25 Jul.–10 Aug. 1990, D.M. Wood] (CNCI C-298).

SLOVAKIA: 1 ♂, 3 ♀♀ [Čachtice environs, 2 Aug. 1991, L. Masner, sweep] (CNCI C-267); Koṧice, 14–17 Sep. 1984, N.D. Springate] (BMNH).

SWEDEN: 1 ♀ [Sk. Röstanga, 6 Jul. 1938, D.M.S.P. and J.F.P.] (BMNH B.M. 1938-414); 1 ♀ [Sk. Kivik, 19 Jul. 1938, D.M.S.P and J.F.P.] (BMNH B.M. 1938-414).

SWITZERLAND: 3 ♂♂, 9 ♀♀ [Dielsdorf, 650 m, 17 Aug. 1984, L. Masner, sweep] (CNCI C-290); 1 ♂ [Jura, Delémont, CABI Lab., 47°22′42″ N, 7°20′30″ E, 7 Jun. 1999, H. Goulet, spruce/beech] (CNCI C-287); 1 ♀ [Solothurn, Weissenstein, 1225 m, 47°15′10″ N, 7°30′00″ E, 17 Jun. 1999, Goulet/White, lush meadow] (CNCI C-285); 3 ♀♀ [Turgau, Unterwasser environs, 1440 m, 4 Aug. 1984, L. Masner, sweeping] (CNCI C-293).

Distribution

Holartic.

Certain records: Andorra (Ferrer-Suay et al. 2011: 358), Colombia (Ferrer-Suay et al. 2012f: 325), Germany (Hartig 1840: 200; Hübner et al. 2002: 507), Japan (Ferrer-Suay et al. 2013i).

Uncertain records: Canada (Vancouver Island) (Andrews 1978: 70), Canada (British Columbia) (Andrews 1978: 56), Canada (Quebec) (Bouchard et al. 1982: 184), France (Kieffer 1902b: 597; De Gaulle 1908: 26), Hungary (Fülöp et al. 2010: 55), Iceland (Andrews 1978: 87), Poland (Kierych 1979b: 15), Romania (Ionescu 1969: 267), Scotland (Cameron 1886: 88), USA (Alaska) (Andrews 1978: 70), USA (Colorado) (Baker 1896: 132,134).

New records: Austria, Czech Republic, Croatia, Japan, Poland, Russia, Sweden and Switzerland.