Galeopsomyia nana Hansson & Hanson 2023, sp.nov.

Galeopsomyia nana Hansson sp.nov.

(Figs 526, 527, 836, 988)

Diagnosis (female). Antenna with a weak constriction between C1 and C2 (Fig. 836); genal carina present; mesoscutellum (Fig. 526) with weak reticulation on median part and a complete median groove, lateral parts with strong reticulation, meshes elongate, submedian grooves distinct, slightly curved and diverging towards posterior part; dorsellum without median carina; propodeum (Fig. 526) with strong reticulation, callus with four setae; gaster (Fig. 526) ovate, 1.7× as long as wide, tergites with strong reticulation, apex of gonoplac blunt (Fig. 527). Female holotype: length of body 1.4mm (paratypes 1.0– 2.2mm).

Scape yellowish-brown, pedicel and flagellum dark brown. Head with frons and vertex black with metallic blue tinges, clypeus dark brown, antennal scrobes black. Thoracic dorsum black with metallic blue tinges. Legs with fore and mid coxae dark brown, hind coxa black with metallic blue tinges; trochanters yellowish-brown; femora with basal half dark brown and apical half yellowish-brown; tibiae yellowish-white; T1–3 yellowish-white, T4 yellowish-brown. Petiole black. Gaster with Gt 1 black with metallic blue tinges, Gt 2-4 golden-purple, Gt5-7 black; gonoplac black.

Antenna with a weak constriction between C1 and C2. Face below level of toruli with weak reticulation, clypeus smooth, frons rugose, antennal scrobes with weak reticulation; with genal carina. Vertex with strong reticulation outside ocellar triangle, inside ocellar triangle with strong reticulation and a transverse carina; with a carina between posterior ocelli in posterior part.

Mesoscutum with strong reticulation, meshes isodiametric on sidelobes, elongate on midlobe; midlobe with a median groove in posterior two-thirds, and with seven scattered adnotaular setae. Mesoscutellum with weak reticulation on median part and a complete median groove, lateral parts with strong reticulation, meshes elongate; submedian grooves distinct, slightly curved and diverging towards posterior part; with eight setae scattered over lateral parts. Dorsellum with weak sculpture, without median carina. Propodeum with a strong median carina that expands anteriorly and posteriorly; with strong reticulation; callus with four setae. Fore and hind coxae with strong reticulation, mid coxa with weak reticulation. Fore wing with three setae on dorsal surface of submarginal vein; speculum closed; costal setal row broken.

Petiole very short, just a narrow band. Gaster ovate; tergites with strong reticulation; apex of gonoplac blunt.

Relative measurements: head length, dorsal view 18; head length, frontal view 28; POL 7.5; OOL 5; lateral ocellus diameter 3.5; head width 40; mouth width 11; malar space 10; eye length 16.5; scape length 14; scape width 3; pedicel+flagellum length 43; pedicel length 5; pedicel width, dorsal view 3; F1 length 6; F1 width 4; F2 length 7; F2 width 4; F3 length 7; F3 width 4; clava length 16.5; clava width 4.5; C3 length 6; spicule length 1.5; mesosoma length 45; mesosoma width 33; midlobe of mesoscutum length 15.5; mesoscutellum length 17; mesoscutellum width 17; median part of mesoscutellum width (measured medially) 7; median part of mesoscutellum, width in anterior part 6.5; median part of mesoscutellum, width in posterior part 8.5; lateral part of mesoscutellum, width (measured medially) 4; dorsellum length 1.5; propodeum length 8; costal cell length 22; costal cell width (measured at widest part) 2; marginal vein length 23; stigmal vein length 8; gaster length 51; gaster width 31; Gt 2 length (measured medially) 7; Gt 4 length (measured medially) 8; Gt 7 length (measured medially) 5; Gt

7 width (measured at base) 11; longest cercal seta length 11; shortest cercal seta length 4.5.

Male. Length of body 1.1–1.4mm.

Antenna (Fig. 988) with scape widest just above the middle; plaque whitish/infuscate and situated in apical part but distinctly below apex; dorso-basal whorls of long setae present on F1–F4 and C1. Gaster short ovate. Otherwise as in female.

Relative measurements: head length, dorsal view 16; head length, frontal view 28.5; head width 38.5; mouth width 12; malar space 10; eye length 16.5; scape length 14; scape width 4.5; plaque length 3; pedicel length 5; pedicel+flagellum length 58; F1 length 4.5; F1 width 4; F2 length 8; F2 width 4; F3 length 9; F3 width 4; F4 length 9.5; F4 width 3.5; clava length 21; clava width 3.5; mesosoma length 47; mesosoma width 33; gaster length 45; gaster width 27; longest subbasal seta on F1, length 30.

Hosts. Unknown.

Distribution. Costa Rica.

Material examined.

Holotype ♀ COSTA RICA, Puntarenas, Reserva Privada Karen Mogensen, 9º52′N, 85º03′W, 305m, 11-12.ii.2018, J.S. Noyes (NHMUK). Paratypes (33♀ 4♂, CNC, MZLU, MZUCR, NHMUK): 5♀ 2♂ with same label data as holotype; following from same locality as holotype but collected 14-15. ii.2005 (1♀), 19-20.ii.2009 (2♀), 17-18.ii.2011 (7♀ 1♂), 23-24.ii.2013 (3♀ 1♂), 19-20.ii.2016 (2♀), 18-19.ii.2019 (1♀); 1♀ COSTA RICA, Puntarenas, San Vito, Las Cruces, 8º47’N, 82º58’W, 1000- 1300m, 19-22.ii.2008, C. Hansson; 3♀ COSTA RICA, Puntarenas, Parque Nacional Piedras Blancas, Estación El Bonito, 8º43’N, 83º13’W, 100m, 13-14.ii.2012, J.S. Noyes; 3♀ COSTA RICA, Puntarenas, Los Charcos de Osa, 8º40′N, 83º30′W, 50m, 18-19.ii.2008, J.S. Noyes; 1♀ COSTA RICA, Puntarenas, Parque Nacional Corcovado, Estación Los Patos, 08°33’N, 83°30’W, 200m, ii.2000, malaise trap & yellow pan trap, J.S. Noyes & J.A. Azofeifa; 1♀ COSTA RICA, Heredia, Estación Biológica La Selva, LN 264463_532850, 100-200m, 30-31.iii.2002, swept, J. Azofeifa; 1♀ COSTA RICA, Limón, Reserva Biológica Hitoy-Cerere, 9º40′N, 53º02′W, 100m, 22-23.ii.2010, J.S. Noyes; 1♀ from same locality as previous but collected 24-26.ii.2008; 1♀ COSTA RICA, Limón, Valle de la Estrella, Reserva Biológica Hitoy Cerere, Sendero Toma de Agua, LN 184300_ 643500, 140m, 8.v.1999, malaise trap, F. Umaña.

Etymology. From the Latin nana = small, referring to small size of this species.