Cyclocephala fasciolata Bates 1888

Cyclocephala fasciolata species group

(Figs. 2A–D; 6E–F; 9A–D; 12C–D; 13E–F; 14A–B; 15C–D; 16C–E; 17C–E; 18D–G; 20D; 24D, E; 25C–D; 27A– F; 32F; 39)

Description. The Cyclocephala fasciolata species group consists of four species and is characterized by following combination of characters. Color black except pronotum with 2 transverse, slightly oblique, reddish-orange bands; body broad, subrectangular (Fig. 2A–D) (C. melane Bates is totally black (Fig. 6E). Head: Frons mostly impunctate, frontoclypeal suture slightly indicated (Fig. 2A–D). Clypeus with punctures dense to sparse, clypeal apex parabolic, slightly reflexed (Fig. 2A–D). Mandible without teeth, inner apex slight concave (Fig. 9A, C); labium densely setose, apex widely notched, paraglossa slightly developed (Fig. 12C–D); maxilla with galea developed, with 6 teeth, lacinia not developed, apex with slender, long setae (Fig. 9B, D). Epipharynx rectangular, densely setose, apex projecting at center (ventral view, Figs. 13E–F; 14A–B). Antenna with 10 antennomeres, club subequal in length to antennomeres 2–7 (slightly shorter in female) Pronotum: Surface with punctures moderate in density, moderately large to large (especially on sides). Base with marginal bead (Fig. 32A). Elytra: Surface with punctacte rows, punctures of striae and intervals similar to those of pronotum; entire surface often coarsely and transversely wrinkled obscuring rows of punctures. Wing vein AA 3 equal length to AA 4 (Fig. 22A–D). Anterior edge of wing distal of apical hinge glabrous (Fig. 22D). Pygidium: Surface usually moderately densely punctate; punctures moderate in size to more usually moderately large, ocellate, setigerous; setae short, tawny, moderately dense (Fig. 24D–E). In lateral view, surface in males evenly convex, weakly convex to nearly flat in females. Legs: Protibia bidentate in males, basal tooth slightly removed from others (Fig. 18D, F–G). Protarsus in male enlarged, tarsomeres 2–4 successively gradually larger, fourth with small tooth on ventral surface, fifth large, curved, with small angulation at base on ventral side (Figs. 17C–E), all tarsomeres longitudinally striate on dorsal face, median claw large, curved, apex cleft (Fig. 15C–E); external edge with some striations (Fig. 16C–E). Metatarsus subequal in length to metatibia. Females with protibia tridentate (Fig. 18E). Venter: Prosternal process moderately long, columnar, apex obliquely flattened in a transverse oval with anterior 2/3 to 4/5 raised into convex “button”. Parameres: As in Fig. 27A–F. Female similar to male, but protarsus simple. Gonocoxite and gonosternite fused (Fig. 32F).

Composition. This group includes the following five species, of which, two species are distributed in Colombia, and are treated below:

Cyclocephala abrelata Ratcliffe & Cave, 2002: 155 [original combination].

Cyclocephala brevipennis Endrődi, 1985: 69 [original combination].

Cyclocephala fasciolata Bates, 1888: 301 [original combination].

Cyclocephala melane Bates, 1888: 310 [original combination].

Cyclocephala santandereana Neita, new species