﻿Notes on twelve species of jumping spiders from Hainan Island, China (Araneae, Salticidae)

﻿Abstract Three new genera and eleven new species are reported from Hainan Island, China. The new genera are Logunattusgen. nov., including L.dufuisp. nov. (♂), and the generotype L.libaiisp. nov. (♂♀), Qiongattusyuanyeaegen. et sp. nov. (♂♀), and Spiralembolusgen. nov., including the generotype S.yinggelingsp. nov. (♂♀), and S.yuisp. nov. (♂♀). Another six new species are Carrhotusqingzhaoaesp. nov. (♂♀), Gedealiangweiisp. nov. (♂♀), Heliophanoidesmoisp. nov. (♂), Indopadillasongisp. nov. (♂♀), Myrmarachnemixiaoqiisp. nov. (♂♀), and Nandiciusshihaitaoisp. nov. (♂♀). The unknown female of the endemic species, Pancoriushainanensis Song & Chai, 1991 is also described for the first time. Diagnostic photos of these species are provided.


Introduction
Hainan, the second-largest Chinese Island, has presented very high species diversity (Tang and Li 2010). As in most Chinese regions, the taxonomic study of jumping spiders from this island began at the end of the 20 th century (Song et al. 1988), but knowledge has rapidly increased in the last more than three decades, with the series of taxonomic studies and biodiversity surveys conducted (Song et al. 1988;Song 1991;Song and Chai 1991;Peng and Kim 1997;Song and Zhu 1998;Zhang and Li 2005;Peng and Li 2006;Guo 2011;Guo et al. 2011a, b;Barrion et al. 2013;Zhou and Li 2013;Xu et al. 2021;Wang and Li 2022b;Yu et al. 2022). To date, at least 123 salticid species including 47 endemics have been recorded from Hainan (Metzner 2023;Wang and Li 2022b;WSC 2023). However, there is no doubt that the true diversity of jumping spiders from this island remains insufficiently known, and numerous new species or newly recorded species will continue being discovered with further taxonomic studies and broad surveys. Moreover, like the worldwide current situation, the taxonomic study of jumping spiders from this island is also plagued by high rates of taxa known from a single-sex and some poorly known species lacking diagnostic drawings (Wang and Li 2022b).
Female ( Fig. 2A 83, 0.98, 1.25, 1.18, 0.63). Carapace similar to that of male, except paler, covered with dense yellow setae and sparse dark, long setae, without white setae on the lateral of thorax, and clypeus. Abdomen oval, dorsum yellow to brown, setose, with yellow, dark markings of setae. Epigyne ( Fig. 2A, B). Wider than long, with arc-shaped basal plate almost as wide as epigyne; copulatory openings posteriorly located, separated from each other less than their width; copulatory ducts thin, paralleled extending anteromedially and followed by ~ 110° curves, and connected to the anterior portions of spermathecae distally; spermathecae irregular, separated from each other less than 1/6 their width.
Distribution. Known only from the type locality in Hainan Province.
Diagnosis. Heliophanoides moi sp. nov. resembles that of Phintella tengchongensis Lei & Peng, 2013 in having similar palp structure, but it can be easily distinguished by the RTA, which is extending anteroventrally, and slightly less than tibia length in retrolateral view (Fig. 5C), vs. extending anteriorly, and ~ 1.5× longer than tibia in P. tengchongensis (Lei and Peng 2013: fig. 8b). It also somewhat resembles that of Echinussa imerinensis Simon, 1901 in the general shape of palpal structure, but it can be easily distinguished by the tapered RTA, which does not extend ventrally beyond the bulb prolateral margin in retrolateral view (Fig. 5C), vs. the RTA narrowest medio-posteriorly, and extends beyond the bulb prolateral margin in E. imerinensis (see the figure in Prószyński 1987).
Description. Male (Fig. 5 1.40, 0.60, 1.13, 1.15, 053). Carapace red-brown, covered with sparse colourful scale-like setae and thin setae; fovea dark red, longitudinal, linear. Chelicerae red-brown, each with two promarginal teeth and one retromarginal tooth. Endites paler than chelicerae and widened distally. Labium red-brown, bearing brown setae at anterior margin. Sternum slightly longer than wide, with straight anterior margin. Legs pale yellow to red-brown, with strongest legs I bearing three and two pairs of ventral spines on the tibiae and metatarsi. Abdomen elongated, dorsum yellow to brown, with alternate yellow and brown transverse bands, covered wholly by scutum; venter pale, with longitudinal, broad, brown band medially. Palp ( Fig. 5A-C). Tibia slightly wider than long in ventral view; RTA slightly less than tibia length, extending anteroventrally, tapered to pointed tip in retrolateral view; DTA flat, almost triangular in ventral view, almost square, and extending anterodorsally in retrolateral view; cymbium ~ 2× longer than wide, setose; bulb elongated, with sub-triangular posterior lobe, medio-retrolaterally located, lamellar tegular bump; embolus strongly sclerotized, short, curved retrolaterally with blunt tip, accompanied by lamellar process more than four times longer than wide, and with arc-shaped out-margin.
Female  Song (1935Song ( -2008, who has made significant contributions to the taxonomy of Hainan jumping spiders; noun (name) in genitive case.
Diagnosis. The male of Indopadilla songi sp. nov. resembles that of I. sabivia Maddison, 2020 in having the bifurcated RTA, but it can be easily distinguished by the RTA has bar-shaped dorsal ramus in retrolateral view (Fig. 6B), vs. tapered, almost triangular dorsal ramus in I. sabivia (Maddison et al. 2020: fig. 118). The female resembles that of I. cuc Wang, Li & Pham, 2023 in having similar epigyne, but it can be distinguished by the following: (1) the epigynal hood is almost square in ventral view (Fig. 7A), vs. almost half-round in I. cuc (Wang et al. 2023: fig. 14A); (2) the AG extends posteriorly, and has slightly enlarged terminus (Fig. 7B), vs. extending towards lateral sides, and without enlarged terminus in I. cuc (Wang et al. 2023: fig. 14B . Carapace dark red, with a pair of white setae behind the PLEs, and arcshaped, red-orange area bearing dense white setae anteriorly on thorax; fovea red-brown, longitudinal. Chelicerae red-brown, each with four promarginal teeth and one retromarginal fissidental tooth with seven cusps. Endites longer than wide, bearing dense brown setae distally on the inner margins. Labium coloured as endites, bearing brown setae on anterior margin. Sternum yellow to red, ~ 1.5× longer than wide. Legs pale to red-brown, legs I strongest, with three and two pairs of ventral spines on tibiae and metatarsi. Abdomen elongated, dorsum green-brown except the lateral sides white, dotted, with two pairs of muscle depressions medially; venter paler than the dorsum, with four, longitudinal, dotted lines. Palp ( Fig. 6A-C). Tibia longer than wide; RTA bifurcated, with strongly sclerotized, apically pointed ventral ramus and bar-shaped dorsal ramus; cymbium ~ 2× longer than wide, covered with pale setae; bulb longer than wide, swollen medio-posteriorly, with posterior lobe extending postero-retrolaterally; embolus long and broad, distally divided into the strongly sclerotized, needle-shaped portion, and the weakly sclerotized, irregular portion.
Distribution. Known only from the type locality in Hainan Island, China.  Etymology. The specific name is a combination of logun, referring to Dr. Dmitri V. Logunov (Manchester, UK), a leading arachnologist in jumping spiders, and attus, meaning jumper. The gender is masculine.
Diagnosis. Logunattus gen. nov. can be recognized as a member of the tribe Euophryini Simon, 1901 by the similarity of habitus and palpal structure to the representative genus of this tribe, Euochin Prószyński, 2018, especially the presence of white setae on the dorsum of palpal tibia and cymbium and the loop of sperm duct inside the tegulum (Maddison 2015). It can be easily recognized by the dagger-axe-shaped RTA. It resembles that of Euochin Prószyński, 2018 in having similar habitus, the presence of white setae on the dorsum of tibia and cymbium of male palp, and large spermathecae, but it can be distinguished by the following: (1) the embolus is straight or curved, vs. coiled in Euochin (Zha et al. 2014: figs 5, 8, 16, 19;Metzner 2023); (2) the RTA is daggeraxe-shaped in retrolateral view, vs. straight in Euochin (Zha et al. 2014: figs 6, 9, 17, 20;Metzner 2023); (3) the chelicera has a single retrolateral tooth, vs. a retromarginal fissidental tooth with several cusps in the generotype and its congeners of Euochin (see the description in Zha et al. 2014); (4) the epigyne has pair of non-transparent atria lack ridges, vs. transparent atria have concomitant lateral ridges in Euochin (Zha et al. 2014: figs 3, 10, 14, 21;Metzner 2023). The genus also somewhat resembles that of Spiralembolus gen. nov. in having similar habitus and copulatory organs, but it can be easily distinguished by the absence of white setae on clypeus, the dagger-axe-shaped RTA, non-spiralled embolus, the presence of median septum, and accessory glands of copulatory ducts, vs. presence of a cluster of white setae on clypeus, RTA non-daggeraxe-shaped, spiralled embolus, the absence of median septum and accessory glands of copulatory ducts in Spiralembolus (Figs 19, 20A-C, G, 21, 22A-C, G).
Description. Small-sized jumping spider. Sexual dimorphism indistinct. Carapace sub-square, covered with dense white and yellow setae, with elevated cephalic region and sloped thorax; fovea longitudinal, dark, linear. Chelicerae yellow-brown, each with two promarginal teeth and one retromarginal tooth. Endites paler than chelicerae, broadened mediodistally. Labium almost linguiform, paler anteriorly. Sternum longer than wide, almost heart-shaped, with straight anterior margin. Legs yellow to dark brown, covered with sparse spines. Abdomen sub-oval, the dorsum yellow to grey-brown, with small anteromedian scutum that only presented in males; venter paler than dorsum, with pair of longitudinal, central dotted lines.
Palp. Tibia short, with ventro-prolateral bump, dagger-axe-shaped RTA, and dense, white setae dorsally; cymbium longer than wide, with dense white setae dorsally on proximal half; bulb elongate-oval, with distinct posterior lobe; embolus originates from antero-prolateral portion of bulb, forming disc at base, and curved or straight medially, with spinous base apophysis or not.
Epigyne. See the description of the generotype. Composition. The genus includes two species, the generotype, and G. dufui sp. nov.
Distribution. Known only from Hainan Island, China.  Diagnosis. Logunattus dufui sp. nov. resembles that of L. libaii sp. nov. in having similar habitus and palpal structure, but it can be easily distinguished by the embolus, which is straight and with spinous proximal apophysis (Fig. 8B), vs. curved and without apophysis in L. libaii (Fig. 9B).
Description. Male (Fig. 8) Chelicerae each with two promarginal teeth and one retromarginal tooth. Endites broadened medio-distally. Labium linguiform. Sternum longer than wide, with straight anterior margin. Legs yellow to brown, with three and two pairs of ventral spines on tibiae and metatarsi I and II, respectively. Abdomen suboval, dorsum pale to brown, covered with sparse white and brown setae, with two pairs of anterior muscle depressions and anteromedian scutum ca. half the abdominal width and length; venter yellow to pale yellow, with pair of longitudinal, dotted lines medially. Palp ( Fig. 8A-C, E). Tibia very short, with ventro-prolateral bump, and dagger-axe-shaped RTA; cymbium ~ 2× longer than wide in ventral view, covered with dense white setae dorso-proximally; bulb elongated, with tapered, downward extending posterior lobe; embolus short and straight, forming disc and with spinous apophysis at base, and blunt apically.
Female  Diagnosis. The male of Logunattus libaii sp. nov. resembles that of L. dufui sp. nov. in general shape of palp, but it can be easily distinguished by the embolus, which is curved medially, and without proximal apophysis in ventral view (Fig. 9B), vs. straight and with proximal apophysis in L. dufui (Fig. 8B). The female of this new species resembles that of Spiralembolus yinggeling sp. nov. in having similar epigyne, but it can be easily distinguished by the presence of median septum, and accessory glands of copulatory ducts (Fig. 10A, B), vs. absent in S. yinggeling (Fig. 20A, B).
Description. Male (Figs 9, 10C, D Carapace red-brown to dark brown, covered with dense setae; fovea dark, longitudinal, barshaped. Chelicerae red-brown to dark brown, each with two promarginal teeth and one retromarginal tooth. Endites dark yellow, with pale distal-inner margins bearing dense brown setae. Labium darker than endites. Sternum coloured as endites, slightly longer than wide, with straight anterior margin, widest medially. Legs yellow to dark brown, with three and two pairs of ventral spines on tibiae and metatarsi I and II, respectively. Abdomen oval, dorsum with two pairs of anterior muscle depressions, indistinct brown or dark brown stripes, and big, irregular pale marking posteriorly, covered by anteromedian scutum; venter pale, with pair Figure 10. Logunattus libaii sp. nov., male holotype and female paratype A epigyne, ventral B vulva, dorsal C holotype habitus, dorsal D ditto, ventral E female paratype habitus, dorsal F holotype carapace, frontal G holotype chelicera, posterior. Scale bars: 0.1 mm (A, B, G); 0.5 mm (C-F). of longitudinal, dotted lines. Palp (Fig. 9A-C). Tibia short, with ventro-prolateral bump, covered with white dorsal setae; RTA dagger-axe-shaped, almost 1.5 times longer than tibia, with pointed tip; cymbium ~ 1.8× longer than wide in ventral view, covered with dorsal white setae at proximal half; bulb elongate-oval, with blunt posterior lobe extending postero-retrolaterally; embolus originates from the antero-prolateral portion of bulb, forming a disc at base, curved towards prolateral side medially and with rather pointed tip directed towards ~ 10:30 o'clock position.
Female (Fig. 10A, B (Fig. 10E) similar to that of male except paler, and without scutum on the dorsum of abdomen. Epigyne (Fig. 10A, B). Wider than long, with pair of shallow atria anteromedially; copulatory openings almost half round, open towards downward, separated by the slightly raised median septum; copulatory ducts curved medially, and touched distally, with sub-triangular proximal accessory glands extending antero-prolaterally; spermathecae oval; fertilization ducts lamellar, extending transversely.
Distribution. Known only from the type locality in Hainan Island, China. Etymology. The specific name is a patronym of Prof. Xiaoqi Mi, who greatly helped us with this research; noun (name) in genitive case.

Genus
Diagnosis. Myrmarachne mixiaoqii sp. nov. can be easily distinguished from other congeners by the presence of a cluster of cymbial macro-setae above the dRTA, and the pair of bag-shaped structures below the epigynal hood, vs. the absence of a cluster of cymbial macro-setae above the dRTA and without similar bag-shaped structures below the epigynal hood in congeners presently known (see Metzner 2023).
Description. Male (Figs 11, 12E   than wide. Legs yellow to brown, with one ventral spine on patellae I, four and two pairs of ventral spines on tibiae I and metatarsi I, respectively. Abdomen elongated, almost gourd-shaped, constricted at anterior 1/3, dorsum yellow to red-brown, covered wholly by scutum; venter pale to brown. Palp (Fig. 11A-D). Tibia wider than long in ventral view; dRTA bifurcated, with platelike ventral ramus and short, digitiform dorsal ramus; cymbium almost oval, bearing cluster of medio-retrolateral macro-setae above the dRTA; bulb almost round, flat, embolus long, coiled more than two coils, with pointed tip reaches the cymbial tip.
Female (Fig. 12A-D (Fig. 12H) similar to that of male except with much shorter chelicerae each with five promarginal and nine or ten retromarginal teeth, without dorsal scutum on abdomen, and with six pairs of ventral spines on tibiae I. Epigyne (Fig. 12A-D). Longer than wide, with tube-shaped epigynal hood fused with pair of bag-shaped structures posteriorly; atria paired, with invert L-shaped lateral ridges; copulatory ducts long, forming complex paths with three distal coils; spermathecae almost spherical, anteriorly located, separated from each other slightly less than their diameter.
Distribution. Only known from the type locality in Hainan Island, China. Etymology. The specific name is after Prof. Haitao Shi, a leading scientist in turtle conservation; noun (name) in genitive case.

Genus
Diagnosis. The male of Nandicius shihaitaoi sp. nov. resembles that of N. proszynskii Wang & Li, 2021 in having very similar habitus and palpal structure, but it can be easily distinguished by the presence of bRTA (Fig. 13B, C), vs. bRTA absent in N. proszynskii (Wang and Li 2021: fig. 10B, C). The female closely resembles that of Tasa koreana (Wesołowska, 1981) in having very similar epigyne, but it can be easily distinguished by the presence of a basal epigynal plate, and the C-shaped copulatory openings (Fig. 14A), vs. the absence of basal epigynal plate and oval copulatory opeings in T. koreana (Suguro and Yahata 2014: fig. 29). The female also resembles that of Madhyattus jabalpurensis Prószyński, 1992 in having a very similar epigyne, but it can be easily distinguished by the distance between the copulatory openings, which is more than half the epigynal width (Fig.  14A), vs. ~ 1/3 the epigynal width in M. jabalpurensis (Prószyński 1992 : fig. 77).  11 (0.73, 0.25, 0.50, 0.38, 0.25). Carapace yellow except the lateral sides of eye field black, covered with white and brown setae, with cluster of white setae at the median of anterior margin, pair of white stripes of setae laterally on cephalic region, and pair of elongate-oval dark spots medially on eye field; fovea indistinct. Chelicerae dark yellow, each with two promarginal teeth and one retromarginal tooth. Endites longer than wide, widened distally, bearing dense setae distally on inner margins. Labium tapered, almost linguiform. Sternum ~ 1.5× longer than wide, with straight anterior margin. Legs pale to dark yellow, with green-brown stripes on the lateral of femora, patellae, and metatarsi II, III, IV; leg I strongest, with slightly enlarged femora and tibiae. Abdomen elongated, dorsum with several transverse chevron patterns medio-posteriorly; venter pale. Palp (Fig. 13A-C). Tibia wider than long, with three apophyses, including the retrolateral one curved inwards distally and blunt apically, the dorsal one almost triangular, and the broad baso-retrolateral one with several small apophyses; cymbium ~ 1.5× longer than wide, with tuberous proximal apophysis; bulb elongated, swollen medio-posteriorly, with antero-retrolateral bump; embolus originates from the antero-prolateral portion of bulb, short and straight, tapered to the rather pointed tip directed towards ~ 2 o'clock position.
Distribution. Known only from the type locality in Hainan Island, China.
Comments. The new species is placed into the genus provisionally because it shares a very similar habitus and palpal structure with the known congener, Nandicius proszynskii Wang & Li, 2021. However, it is inconsistent with other congeners in the epigynal structure and body shape, which indicates its generic position needs further confirmation.

Genus Pancorius Simon, 1902
Pancorius hainanensis Song & Chai, 1991 Figs 15, 16 Pancorius hainanensis Song & Chai, 1991: 20, fig. 10A, B (male holotype, examined). Diagnosis. The male of Pancorius hainanensis Song & Chai, 1991 resembles that of P. wesolowskae Wang & Wang, 2020 in the general shape of palpal structure, especially the small, blunt posterior lobe, but it can be easily distinguished by the following: (1) the RTA being slightly greater than sperm duct diameter in width, and with a pointed tip in retrolateral view (Fig. 15B), vs. more than two times greater than sperm duct diameter in width, and blunt apically in P. wesolowskae (Wang and Wang 2020: fig. 20), (2) the embolus acutely narrowed to the pointed tip distally in ventral view (Fig. 15A) vs. almost tapered in P. wesolowskae (Wang and Wang 2020: fig. 19). The female of this species resembles that of P. crinitus Logunov & Jäger, 2015 in having very shallow epigynal hoods, but it can be easily distinguished by anterior chamber of spermathecae, which are wider than long (Fig. 16B), vs. ~ 2× longer than wide in P. crinitus (Logunov and Jäger 2015: fig. 43).

Type material examined. Holotype
Description. Male (Figs 15, 16C, D 2.50, 1.13, 1.80, 1.85, 0.88). Carapace red-brown to dark brown, covered with white and dark setae, dense on both sides; fovea longitudinal, dark. Chelicerae red-brown, each with two promarginal teeth and one retromarginal tooth. Endites paler than chelicerae, bearing dense dark brown setae on the distal half of inner margins. Labium tapered, with pale distal end. Sternum yellow to red-brown, shield-shaped, ~ 1.5× longer than wide. Legs yellow to dark, setose, and spiny. Abdomen elongated, dorsum dark brown and mingled with green, spotted, with longitudinal, pale band anteromedially, two pairs of median muscle depressions followed by four transverse arc-shaped pale stripes; venter dark brown, spotted laterally, with pair of longitudinal, dotted lines medially. Palp (Fig. 15A-C). Tibia longer than wide, with strongly sclerotized RTA slightly curved inward distally and pointed apically; cymbium 1.5× longer than wide; bulb slightly swollen, with small bean-shaped posterior lobe; embolus short, broad, and flat, slightly curved retrolaterally, with pointed tip.
Female (Fig. 16A, B (Fig. 16E) similar to that of male except without dense setae on the carapace and with an almost T-shaped yellow area on thorax. Epigyne (Fig. 16A, B). Wider than long, with pair of very shallow posterior hoods; copulatory openings slit-shaped,  Diagnosis. The male of Qiongattus yuanyeae sp. nov. resembles that of Chinattus chichila Logunov, 2003 in the general shape of palp, but it can be easily distinguished by the narrow cymbium, which is ~ 3× longer than wide in ventral view (Fig. 17B), vs. less than 2× longer than wide in C. chichila (Logunov 2003: fig. 1), and by the RTA, which is acutely narrowed distally in retrolateral view (Fig. 17C), vs. tapered in C. chichila (Logunov 2003 : fig. 2). The female of this species resembles that of Chinattus inflatus Wang & Li, 2020 in having Figure 18. Qiongattus yuanyeae sp. nov., male holotype and female paratype A, B epigyne, ventral C vulva, dorsal D holotype habitus, dorsal E ditto, ventral F female paratype habitus, dorsal G holotype carapace, frontal H holotype chelicera, posterior. Scale bars: 0.1 mm (A-C, H); 0.5 mm (D-G). similar basal epigynal plate, but it can be easily distinguished by the presence of spermathecal head and the absence of round structure on basal epigynal plate ( Fig. 18A-C) vs. spermathecae lack heads and basal epigynal plate has round structure in C. inflatus (Wang and Li 2021: fig. 1A, B).
Distribution. Known only from the type locality in Hainan Island, China. Etymology. The specific name is a combination of spiral and embolus, referring to the species that has spiral embolus. The gender is masculine.

Genus
Diagnosis. Spiralembolus gen. nov. can be recognized as a member of the tribe Euophryini Simon, 1901 for the same reasons as Logunattus gen. nov. It is remarkable for the thick, spiral embolus. It above diagnosed with Logunattus gen. nov. It also resembles that of Euochin Prószyński, 2018 in having similar habitus and palpal structure, especially the presence of white setae on the dorsum of palpal tibia and cymbium, and the oval or round spermathecae, but it can be easily distinguished by the following: (1) the thick, spiralled embolus, not forming a disc at base, vs. flagelliform, coiled embolus, and mostly forming a disc at base in Euochin (Zha et al. 2014: figs 5, 8, 16, 19;Metzner 2023); (2) the presence of a cluster of white setae on clypeus, and dorsal abdominal scutum in males, vs. absent in Euochin (Zha et al. 2014: figs 2, 13;Metzner 2023); (3) the epigyne lacking pair of oval or round transparent atria, and the concomitant lateral ridges, vs. present in Euochin (Zha et al. 2014: figs 3, 10, 14, 21;Metzner 2023). The genus also somewhat resembles that of Chalcovietnamicus Marusik, 1991 in having similar epigyne, but it can be easily distinguished by the presence of clusters of white setae on clypeus, and the dorsum of tibia and cymbium of palp, the spiralled embolus lacks lamellar basal apophysis, vs. without clusters of white setae on clypeus, and the dorsum of tibia and cymbium of palp, non-spiralled embolus with lamellar basal apophysis in Chalcovietnamicus (Żabka 1985: figs 71-74;Wang and Li 2022a: figs 3A-C, 4F).
Description. Small-sized jumping spider. Sexual dimorphism scarcely evident. Carapace darker in males and, covered with cluster of white setae on clypeus that only presents in males, with elevated, sub-square cephalic region bearing thin setae, and sloped thorax with dark brown stripes; fovea dark red, longitudinal. Chelicerae each with two promarginal teeth and one retromarginal tooth. Endites widened distally, with paler inner margins bearing dense brown setae. Labium tapered, almost linguiform. Sternum longer than wide, with straight anterior margin, bearing brown and pale setae of various lengths. Legs yellow to dark brown, spinous, with clusters of white setae on femora and tibiae that only present in males. Abdomen darker in males, dorsum covered with dense white setae and with antero-median scutum, those only present in males, and with several transverse, pale stripes posteriorly in both sexes; venter setose.
Palp. Tibia very short, covered with dorsal white setae; RTA short, curved medially or distally, with rather blunt tip; cymbium longer than wide, covered with dorsal white setae on proximal half; bulb swollen; embolus thick, strongly sclerotized, originates from the prolatero-apical portion of bulb, spiralled into coils.
Epigyne. Wider than long, without distinct atrium; copulatory openings located anteriorly or medially, round or oval, separated from each other at least their width; copulatory ducts short, strongly curved anteriorly; spermathecae oval or spherical, with distinct Bennett's glands; fertilization ducts originate from the anterior portions of spermathecae, extending transversely.
Composition Etymology. The specific name is after the holotype locality, Yinggeling National Nature Reserve ; noun (name) in apposition.
Description. Male (Figs 19, 20D with cluster of white setae on clypeus, and oblique, dark brown stripes on thorax; fovea longitudinal, dark, linear. Chelicerae dark yellow to red-brown, each with two promarginal teeth and one retromarginal tooth. Endites almost square, with paler inner margins bearing brown setae. Labium tapered, bearing several dark setae on paler distal portion. Sternum almost oval, yellow-brown, and covered with dark setae. Legs yellow to dark brown, bearing dense dark brown ventral setae on tibiae I and clusters of white setae on femora and tibiae. Abdomen oval, dorsum dark brown, covered with white and dark setae and anteromedian scutum, with irregular pale-yellow patches medially and transverse, pale stripes posteriorly; venter dark brown, covered with short dark brown setae, with pair of mediolateral pale patches. Palp (Figs 19A-C, 20E). Tibia short, covered with dense white, long setae dorsally, with tapered RTA curved towards ventral side medially and blunt apically; cymbium ~ two times longer than wide, covered with dense, long, white setae on the dorsum of proximal half; bulb swollen; sperm duct strongly curved retrolaterally; embolus originates from the antero-prolateral portion of bulb, spiralled into less than three coils, with rather pointed tip.
Female ( Fig. 20A- (Fig. 20F) similar to that of male except paler, without dense dorsal white and dark setae, and the dorsal scutum on the abdomen, the cluster of white setae on clypeus, and clusters of white setae on femora and tibiae of legs. Epigyne (Fig. 20A-C). Wider than long; copulatory openings round, anteromedially located, separated from each other ~ 1.5× equal to their width; copulatory ducts thick, inner to spermathecae, close to each other, strongly curved anteromedially, and connected to the posterior portions of spermathecae; spermathecae almost oval, ~ 2× wider than copulatory ducts; fertilization ducts originate from the most antero-inner portions of spermathecae, transversely extending.
Distribution. Known only from the type locality in Hainan Island, China.  , one of the famous ancient Chinese poets and one sorrowful ancient Chinese Emperor; noun (name) in genitive case.
Distribution. Known only from the type locality in Hainan Island, China.