﻿Revision of the genus Colasia Koch, 1965 (= Belousovia Medvedev, 2007, syn. nov.) (Coleoptera, Tenebrionidae, Blaptini)

﻿Abstract The relationship between the genera Colasia Koch, 1965 and Belousovia Medvedev, 2007 within the tribe Blaptini is discussed, and a new synonymy is proposed: Belousovia Medvedev, 2007, syn. nov. of Colasia Koch, 1965. As a result, three new combinations are established: Colasiahelenae (Medvedev, 2007), comb. nov., C.kabakiintermedia (Medvedev, 2007), comb. nov., and C.kabakikabaki (Medvedev, 2007), comb. nov.Colasiaakisoides Koch, 1965 is redescribed, and a lectotype is designated. Three new species of the genus Colasia are described and illustrated from China: C.bijicasp. nov. (Guizhou), C.medvedevisp. nov. (Yunnan), and C.pilosasp. nov. (Yunnan). A distribution map and a key to species of the revised genus Colasia are presented.

The genus Belousovia, named for Igor Alexandrowich Belousov, is represented by one species and two subspecies described from western Yunnan, China: B. helenae Medvedev, 2007, B. kabaki intermedia Medvedev, 2007, andB. kabaki kabaki Medvedev, 2007. According to the original description , Belousovia closely resembles Colasia (based on the examination of C. medvedevi sp. nov., erroneously determined by N. Skopin and G. Medvedev as C. akisioides) in the structure of aedeagus, spiculum gastrale, ovipositor, and elytra, but clearly differs from Colasia by the structure of legs, female genital tubes, and head capsule. The first and the most specific character, male legs: in Belousovia species, the ventral surface of pro-and mesotarsomeres I-IV, and metatarsomeres I-III with long and dense hairy tuft, the apical part of the metatibiae with a row of dense setae on the inner side (e.g., B. kabaki kabaki Medvedev, 2007: figs 116, 117); in Colasia species, the ventral surface of tarsal segments without hairy brush or tuft (from , only very short and strong setae present, metatibiae without a row of setae on the inner side. The second character, head capsule: the labrum and apical maxillary palpomere are covered with long setae in Belousovia species, but are short in Colasia species; eyes short and distinctly arcuately projecting outwards in Belousovia species, but absolutely flat in Colasia species. The third character, appearance: the pronotum is obviously heart-shaped in Colasia species, which never occurs in Belousovia species; the body is coal-black in Colasia species, but appears reddish in strong illumination in Belousovia species, especially on the humeral carinae and declivity of the elytra. The last character, female genital tubes: in Colasia species, bases of first and second reservoirs and base of spermathecal sphincter diverge from one point; in Belousovia species, the base of the first reservoir is separated from the place of divergence of the bases of the second reservoir and spermathecal sphincter by a very long duct. However, the boundary between the genera Belousovia and Colasia becomes blurred with the examination of additional materials from western Guizhou, and eastern, central and western Yunnan, China.
This study aims to investigate the taxonomic status of the genera Colasia and Belousovia. Additionally, a redescription of C. akisoides and descriptions of three new Chinese species are provided.

Material and method
The specimens were examined and dissected under a Nikon SMZ800 microscope, and photographs were taken using Canon EOS 5DSR camera and processed by Adobe Photoshop 2021. The distribution map was made by QGIS and processed by Adobe Photoshop 2021. Aedeagi was detached from the body with insect pins, then glued to separate cards and pinned under the specimens. A single slash (/) separates data of different lines on a label, a double slash (//) separates data of different labels, authors' remarks are enclosed in brackets "[ ]".
Specimens examined in this study are deposited at the following institutes and collections:

Remarks.
After the examination of types of the genera Belousovia and Colasia, and also additional materials, we propose the genus Belousovia Medvedev, 2007 as a junior synonym of the genus Colasia  Firstly, in male legs: after the re-examination of the types of Colasia akisoides, apical part of metatibiae with a row of setae on the inner side (Fig. 5G, G'; not mentioned in the original description by , although less dense than Belousovia species (Fig. 19G, G'), but dense enough compared to materials from Western Guizhou, and Western Yunnan (Figs 6G, G', 20; a few setae present only); ventral surface of tarsal segments with hairy tuft at apex (Fig. 5E, F, H). As emphasized by , the degree of development of the setae on ventral surface of pro-and mesotarsomeres of the male, and the setae on the inner side of tibiae steadily characterizes morphological distinctiveness of separate genera in the subtribe Gnaptorinina. However, Medvedev is sometimes a little over-dependent on the degree of development of the setae (Medvedev 2009), and this has been questioned by the molecular evidence . Moreover, acquired activities can also damage setae. Thus, the first and the most important evidence presented by  is untenable. Secondly, in head capsule, and in appearance: setae on labrum and maxillary palpi, size and shape of eyes, shape of pronotum, and coloration of body are common characters, they may be distinct in separate genera, but are also variable between different species within a genus. Besides, the author is a bit exaggerates the differences between the genera Belousovia and Colasia, as the pictures presented in the publication . Lastly, the female genital tubes can indeed be used to separate genera, as the author used in the diagnosis (Medvedev 2007), but is not a decisive character. Therefore, the last three distinctions observed by Medvedev are perhaps not enough to distinguish the genera Belousovia and Colasia. Koch, 1965 Figs 1-5, 28, 29, 33 Colasia akisoides Koch, 1965: 131;Medvedev 2001: 95;Löbl et al. 2008   Remarks. This species was described based on the collection of Muséum national d'Histoire naturelle, Paris, France , including ten specimens from China in the original description: "2♂♂ 2♀♀ <<Giufu-Shan, Szechuan>>! leg. Emm. REIT-TER; 1♂ 4♀♀, Kintushan, Prov. Szechuan, West-China>>! leg. BECKER; 1 ♀ <<Junan, Junan-fu>>!".

Colasia akisoides
In January 2017, the first author had a chance to visit the Muséum national d'Histoire naturelle, but no types of this species have been found. In May 2022, XB asked Dr. Antoine Mantilleri (curator for Coleoptera at the MNHN) for help in searching for the types of this species. No such specimens were identified. Fortunately, three syntypes (1♂, 2♀) of this species were founded in HNHM and lent us to study by late Dr. Ottó Merkl. Therefore, the male type deposited in HNHM is designated as the lectotype in this paper, and the remaining types becoming paralectotypes.
The identification of the female paralectotype from Junan-fu (most likely Kunming City), Junan (Yunnan) is in doubt. On the one hand, according to the distribution of all the species of the revised genus Colasia, it is unlikely to be C. akisoides. On the other hand, it was once identified as Asidoblaps glyptoptera by Gebien . Recently, we have obtained some specimens from Kunming, Yunnan, and they were confirmed to be Asidoblaps sp. after the identification. These specimens superficially resembled representatives of Colasia, but were obviously different by aedeagal morphology. Thus, we speculate that the female paralectotype from Yunnan belongs to the Asidoblaps species.
Giufu-Shan and Kinfushan (sometimes erroneously spelled Kintushan) both refer to the current Jinfo Shan (Bezděk et al. 2015;Kataev and Liang 2015;Ge et al. 2021). Jinfo Shan is located in southern Chongqing, Chongqing once belonged to Sichuan Province, and now it is a municipality directly under the Central Government. Thus, the type locality of this species is Jinfo Shan, and the distribution of this species should be changed from Sichuan and Yunnan (Löbl et al. 2008;Ren et al. 2016;Nabozhenko and Chigray 2020) to Chongqing.
Redescription. Body black, weakly shiny; legs shiny. Male. Head. Apical maxillary palpomere triangular, covered with moderately dense and long setae. Anterior margin of labrum emarginate, lateral margins weakly arcuate. Anterior margin of epistoma emarginate; surface flat, matte, inconspicuous punctate. Frontoepistomal suture shallow and arcuate. Dorsal surface of head flat, matte, sparsely and finely granulated. Genal margins arcuately converging forwards, densely and shallowly punctures merged into short wrinkles. Emargination of outer margins of head above antennal base straight. Eyes transverse, not protruding beyond contour of head, distance between outer margins of eyes represent the widest of head; height 0.53 mm, width 0.17 mm from lateral view, respectively (height 3.1× width; 2.5× if the height and width rounded to one decimal place). Temples arcuately narrowing backwards, weakly granulated. Antennae slender and long, with the last segment reaching beyond pronotal base; basal part of antennomere I invisible in dorsal view; II-VII cylindrical, thicker at apex, II very short, III very long, V-VI equal in length, longer than IV and shorter than VII; VIII-X nearly spherical; XI sharped-oval.
Prothorax. Pronotum cordiform, widest at middle, 1.5× wide as long, 1.8× wide as head, ratio of width at anterior margin to middle and base 7: 11: 8; anterior margin deeply emarginate, beaded laterally; lateral margins weakly "S" curved, entirely beaded and smooth; posterior margin straight at middle, beaded laterally; anterior angles rectangular and protruding forwards, posterior angles rectangular; surface matte, central convex, lateral sides weakly depressed along lateral margins, with shallowly and rounded depressions in sides of central, moderately depressed near posterior angles in sides of base, longitudinal median line (median depression in Medvedev 2007) smooth and inconspicuous; shallowly, sparsely and finely punctate in central part, near posterior margin, lateral margins, and lateral sides of anterior margin with wrinkly punctures, sparsely and finely granulated. Prothoracic hypomera depressed, densely and shallowly wrinkled in longitudinal, with sparse and tiny granules. Prosternal process sharply sloping downwards behind procoxae, apex blunt in lateral view.
Pterothorax. Elytra oval, widest at middle, 1.3× long as wide, 1.3× wide as pronotum; base nearly as wide as pronotal base; dorsal surface matte, weakly convex, declivity sharply sloping downwards; humeral carinae smooth, with very sparse, smooth and large tubercles, sparse and fine granules, sparse, coarse and shallow wrinkles between humeral carinae; each elytron with two carinae (sensu Medvedev 2007) between suture and humeral carina, the second carina inconspicuous; declivity with sparse and short setae, slightly convex along suture; surface of epipleuron (sensu Kamiński et al. 2019) matte, edge relatively wide; pseudopleuron (outer part of elytra in Medvedev 2007) much wider than epipleuron, surface matte, with sparse and inconspicuous granules, edge thin and elevated, reaching sutural angle. Scutellum triangular, covered by pronotum.
Legs. Slender and long. Femora claviform, mesofemora slightly longer than profemora, and shorter than metafemora. Protibiae weakly curved, distal part distinctly thick; mesotibiae and metatibiae straight, both gradually widened toward apex; distal part of metatibiae with a row of golden yellow hairy row on inner side. Ventral surface of proand mesotarsomeres I-IV, and metatarsomeres I-III with undeveloped hairy tuft at apex.
Aedeagus. Length 1.8 mm, width 0.5 mm. Parameres length 0.5 mm, width 0.4 mm. Parameres relatively wide and short, widest at base, basal 1/4 parallel, and then narrowing toward apex nearly straight, distal part weakly curved to ventral side in lateral view.
Female. Antennae not reaching pronotal base; elytra wider and more convex; abdominal ventrites 1 and 2 convex; inner side of metatibiae without golden yellow setae; other characters similar to male.
Measurements. Body length: 11.5-13.5 mm; width: ♂ 5.5-6.0 mm, ♀ 6.5-7.0 mm. Diagnosis. This new species closely resembles C. akisoides based on the pronotum cordiform, but can be distinguished from the latter by the following characters (based on male): (1) emargination of outer margins of head above antennal base widely obtuse-angular (straight in C. akisoides); (2) distal part of metatibiae with a few golden yellow spines on inner side (with a row of golden yellow hairy row in C. akisoides); (3) parameres relatively narrow and long (wide and short in C. akisoides), widest at base, and narrowing toward apex nearly straight (basal 1/4 parallel, and then narrowing toward apex nearly straight in C. akisoides), distal part nearly straight in lateral view (weakly curved to ventral side in C. akisoides). This new species is also somewhat similar to C. medvedevi sp. nov. based on the distal part of metatibiae with a few golden yellow setae on inner side in male, it differs from the later by the following characters (based on male): (1) eyes not protruding beyond contour of head (slightly protruding beyond contour of head in C. medvedevi sp. nov.); (2) pronotum cordiform (transverse, subcordiform in C. medvedevi sp. nov.), anterior and posterior angles rectangular (nearly rectangular in C. medvedevi sp. nov.); (3) distal part of protibiae distinctly thick (protibiae gradually widened toward apex in C. medvedevi sp. nov.); (4) parameres relatively narrow and long (wide and short in C. medvedevi sp. nov.), widest at base, and narrowing toward apex nearly straight (basal 1/3 parallel, and then narrowing toward apex nearly straight in C. medvedevi sp. nov.), distal part nearly straight in lateral view (weakly curved to ventral side in C. medvedevi sp. nov.).
Distribution. China: Guizhou. Etymology. The species name is derived from the type locality Bijie. Description. Body black, weakly shiny; legs shiny. Male. Head. Apical maxillary palpomere triangular, covered with moderately dense and long setae. Anterior margin of labrum emarginate, lateral margins parallel. Anterior margin of epistoma emarginate, nearly straight at middle; surface flat, matte, inconspicuous punctured. Frontoepistomal suture shallow and arcuate. Dorsal surface of head flat, matte, sparsely and finely granulated. Genal margins arcuately converging forwards, densely and shallowly punctures merging into shallow wrinkles. Emargination of outer margins of head above antennal base widely obtuse-angular. Eyes transverse, not protruding beyond contour of head, distance between outer margins of eyes represent the widest of head; height 0.53 mm, width 0.17 mm from lateral view, respectively (height 3.1× width; 2.5× if the height and width rounded to one decimal place). Temples arcuately narrowing backwards, weakly granulated. Antennae slender and long, with the last segment reaching beyond pronotal base; basal part of antennomere I invisible in dorsal view; antennomeres II-VII cylindrical, thicker at apex, II shortest, III longest, V-VI equal in length, longer than IV and shorter than VII; antennomeres VIII-X nearly spherical; antennomere XI sharped-oval.
Prothorax. Pronotum cordiform, widest at middle, 1.4× wide as long, 1.8× wide as head, ratio of width at anterior margin to middle and base 8: 12: 10; anterior margin deeply emarginate, beaded laterally; lateral margins weakly "S" curved, entirely beaded and smooth; posterior margin bisinuate, beaded laterally; anterior angles rectangular and protruding forwards, posterior angles rectangular; surface matte, central convex, lateral sides weakly depressed along lateral margins, with shallowly and rounded depressions in sides of central, moderately depressed near posterior angles in sides of base, longitudinal median line smooth and inconspicuous; shallowly, sparsely, and finely punctate in cen-tral part, near posterior margin, lateral margins, and lateral sides of anterior margin with wrinkly punctures, sparsely and finely granulated. Prothoracic hypomera depressed, densely and shallowly wrinkled in longitudinal, with sparse and tiny granules. Prosternal process sharply sloping downwards behind procoxae, apex blunt in lateral view.
Pterothorax. Elytra oval, widest near middle, 1.3× long as wide, 1.4× wide as pronotum; base nearly as wide as pronotal base; dorsal surface matte, weakly convex, declivity sharply sloping downwards; humeral carinae smooth, with very sparse, smooth and large tubercles, sparse and fine granules, sparse, coarse and shallow wrinkles between humeral carinae; each elytron with two carinae between suture and humeral carina, the second carina inconspicuous; declivity with sparse and short setae, slightly convex along suture; surface of epipleuron matte, edge relatively wide; pseudopleuron much wider than epipleuron, surface matte, with sparse and inconspicuous granules, edge thin and elevated, reaching sutural angle. Scutellum triangular, covered by pronotum.
Legs. Slender and long. Femora claviform, mesofemora slightly longer than profemora, and shorter than metafemora. Protibiae weakly curved, distal part distinctly thick; mesotibiae nearly straight, metatibiae straight, both gradually widened toward apex; distal part of metatibiae with a few golden yellow setae on inner side. Ventral surface of proand mesotarsomeres I-IV, and metatarsomeres I-III with undeveloped hairy tuft at apex.
Aedeagus. Length 1.7 mm, width 0.5 mm. Parameres length 0.6 mm, width 0.4 mm. Parameres relatively narrow and long, widest at base, and narrowing toward apex nearly straight, distal part nearly straight in lateral view.
Distribution. China: Yunnan. Etymology. The species name is derived from the name of Prof. Gleb Sergeevich Medvedev, in memory of his outstanding contribution to the knowledge of the tribe Blaptini of the tenebrionid beetles.
Description. Body black, weakly shiny; legs shiny. Male. Head. Apical maxillary palpomere triangular, covered with moderately dense and long setae. Anterior margin of labrum slightly emarginate, lateral margins parallel. Anterior margin of epistoma slightly emarginate; surface flat, matte, shallowly punctate. Frontoepistomal suture shallow and arcuate. Dorsal surface of head flat, matte, sparsely and finely granulated. Genal margins arcuately converging forwards, densely and shallowly punctures merged into short wrinkles. Emargination of outer margins of head above antennal base widely obtuse-angular. Eyes transverse, slightly protruding beyond contour of head, distance between outer margins of eyes represent the widest of head; height 0.52 mm, width 0.19 mm from lateral view, respectively (height 2.7× width; 2.5× if the height and width are rounded to one decimal place). Temples arcuately narrowing backwards, sparsely granulated. Antennae slender and long, with the last segment reaching beyond pronotal base; basal part of antennomere I invisible in dorsal view; antennomeres II-VII cylindrical, thicker at apex, II very short, III very long, V-VI equal in length, slightly longer than IV and shorter than VII; VIII-X nearly spherical; XI sharped-oval.
Prothorax. Pronotum transverse, subcordiform, widest at middle, 1.5× wide as long, 1.9× wide as head, ratio of width at anterior margin to middle and base 7: 11: 8; anterior margin deeply emarginate, beaded laterally; lateral margins weakly "S" curved, entirely beaded and smooth; posterior margin straight at middle, beaded laterally; anterior angles nearly rectangular and protruding forwards, posterior angles nearly rectangular; surface matte, central convex, lateral sides weakly depressed along lateral margins, with shallowly and rounded depressions in sides of central, moderately depressed near posterior angles in sides of base, longitudinal median line smooth and weak; shallowly, sparsely, and finely punctate in central part, near posterior margin, lateral margins, and lateral sides of anterior margin with wrinkly punctures, sparsely and finely granulated. Prothoracic hypomera depressed, densely and shallowly wrinkled in longitudinal, with sparse and tiny granules. Prosternal process sharply sloping downwards behind procoxae, apex blunt in lateral view.
Pterothorax. Elytra oval, widest at middle, 1.3× long as wide, 1.3× wide as pronotum; base nearly as wide as pronotal base; dorsal surface matte, relatively flat, declivity sharply sloping downwards; humeral carinae smooth, with very sparse, smooth and large tubercles, sparse and fine granules, sparse, coarse and shallow wrinkles between Figure 19. Characters of Colasia kabaki kabaki (Medvedev) (male) A head B antenna C pronotum D protibia E protarsus F mesotarsus G metatibia G' distal part of metatibia H metatarsus I, J aedeagus in dorsal and lateral view, respectively. Scale bars: 1.0 mm. humeral carinae; each elytron with two carinae between suture and humeral carina, the second carina inconspicuous; declivity with sparse and short setae; surface of epipleuron matte, edge relatively wide; pseudopleuron much wider than epipleuron, surface matte, with sparse and inconspicuous granules, edge thin and elevated, reaching sutural angle. Scutellum triangular, covered by pronotum. Abdomen. Apex of ventrite 1, and base of ventrite 2 flat in middle; ventrites 1, 2, and anterior part of ventrite 3 rough, with moderately dense and long setae, and inconspicuous granules, densely, shallowly, and finely wrinkled at sides; posterior part of ventrite 3 smooth, shallowly punctate; ventrite 4 smooth, shallowly and finely punctate; last ventrite smooth, with sparse, fine punctures and short setae, apical margin widely rounded.
Legs. Slender and long. Femora claviform, mesofemora slightly longer than profemora, and shorter than metafemora. Protibiae weakly curved, mesotibiae nearly straight, metatibiae straight, both gradually widened toward apex; distal part of metatibiae with a few golden yellow setae on inner side. Ventral surface of pro-and mesotarsomeres I-IV, and metatarsomeres I-III with undeveloped hairy tuft at apex.
Aedeagus. Length 1.3 mm, width 0.6 mm. Parameres length 0.7 mm, width 0.5 mm. Parameres relatively wide and short, widest at base, basal 1/3 parallel, and then narrowing toward apex nearly straight, distal part weakly curved to ventral side in lateral view.
Female. Antennae not reaching pronotal base; elytra wider and convex; abdominal ventrites 1 and 2 convex; inner side of metatibiae without golden yellow setae; other characters similar to male.
Measurements. Body length: ♂ 12.8-13.5 mm, ♀ 13.0-14.0 mm; width: ♂ 6.2-6.5 mm, ♀ 7.2-7.5 mm.   Diagnosis. This new species is obviously different from other species in the following characters (based on male): body large size, length 15.5-17.5 mm (vs. 11.1-13.8 mm in other species); inner side of metatibiae with a row of well-developed densely hairy brush from middle to apex; ventral surface of pro-and mesotarsomeres I-IV, and metatarsomeres I-III with well-developed hairy brush from middle to apex; parameres laminar in dorsal view.
Distribution. China: Yunnan. Etymology. The species name is derived from the dense hairy brush of male metatibiae. Description. Body black, weakly shiny. Male. Head. Apical maxillary palpomere triangular, covered with moderately dense and long setae. Anterior margin of labrum slightly emarginate, lateral margins weakly arcuate. Anterior margin of epistoma slightly emarginate; surface flat, matte, shallowly punctate. Frontoepistomal suture shallow and arcuate. Dorsal surface of head flat, matte, sparsely granulated at sides. Genal margins arcuately converging forwards, densely and shallowly punctate. Emargination of outer margins of head above antennal base widely obtuse-angular. Eyes transverse, protruding beyond contour of head, distance between outer margins of eyes represent the widest of head; height 0.80 mm, width 0.26 mm from lateral view, respectively (height 3.1× width; 2.7× if the height and width are rounded to one decimal place). Temples arcuately narrowing backwards, sparsely granulated. Antennae slender and long, with the last segment reaching beyond pronotal base; basal part of antennomere I invisible in dorsal view; antennomeres II-VII cylindrical, thicker at apex, II very short, III very long, IV and VI subequal in length, V and VII subequal in length; VIII-X nearly spherical; XI sharped-oval.
Prothorax. Pronotum transverse, subcordiform, widest at middle, 1.5× wide as long, 1.9× wide as head, ratio of width at anterior margin to middle and base 6: 10: 8; anterior margin deeply emarginate, beaded laterally; lateral margins weakly "S" curved, entirely beaded and smooth; posterior margin straight at middle, beaded laterally; anterior angles nearly rectangular and protruding forwards, posterior angles nearly rectangular; surface matte, central convex, lateral sides weakly depressed along lateral margins, with moderately depressions in middle of sides and near posterior angles in sides of base, sometimes shallowly and transversely depressed before posterior margin in middle, longitudinal median line inconspicuous; shallowly, sparsely, and finely punctate in central part, near posterior margin, lateral margins, and anterior margin with wrinkly punctures, sparsely and finely granulated. Prothoracic hypomera depressed, densely and shallowly wrinkled in longitudinal, with sparse and tiny granules. Prosternal process sharply sloping downwards behind procoxae, apex blunt in lateral view.
Pterothorax. Elytra oval, widest at middle, 1.2× long as wide, 1.4× wide as pronotum; base wider than pronotal base; dorsal surface matte, relatively flat, declivity sharply sloping downwards; humeral carinae smooth, with very sparse, smooth and large tubercles, sparse and fine granules, sparse, coarse and shallow wrinkles between humeral carinae; each elytron with two carinae between suture and humeral carina, the second carina inconspicuous; declivity with sparse and short setae, slightly convex along suture; surface of epipleuron matte, edge relatively wide; pseudopleuron much wider than epipleuron, surface matte, sparsely granulated, edge thin, reaching sutural angle. Scutellum triangular, covered by pronotum.
Legs. Slender and long. Femora claviform, mesofemora slightly longer than profemora, and shorter than metafemora. Protibiae and metatibiae weakly curved, mesotibiae straight, both gradually widened toward apex; inner side of metatibiae with a row of densely golden yellow hairy row from middle to apex. Ventral surface of pro-and mesotarsomeres I-IV, and metatarsomeres I-III with dense hairy brush from middle to apex.
Aedeagus. Length 2.8 mm, width 0.9 mm. Parameres length 1.0 mm, width 0.7 mm. Parameres relatively wide and short, widest at base, and narrowing toward apex nearly straight, distal part straight in lateral view.
Female. Antennae not reaching pronotal base; elytra wider and more convex; abdominal ventrites 1 and 2 convex; inner side of metatibiae without golden yellow hairy row; ventral surface of tarsi without hairy brush; other characters similar to male.