Gary R. Graves,¹ Gerald J. Niemi<sup>2

1</sup>Department of Vertebrate Zoology, MRC-116, National Museum of Natural History, Smithsonian Institution, P.O. Box 37012, Washington, D.C. 20013-7012, U.S.A. gravesg@si.edu
²Department of Biology and the Center for Water and the Environment, Natural Resources Research Institute, University of Minnesota, 5013 Miller Trunk Highway, Duluth, MN 55811-1442, U.S.A