New pseudoscorpion records (Arachnida: Pseudoscorpiones) from Lorestan province, western Iran, with redescriptions of Olpium lindbergi (Olpiidae) and Geogarypus shulovi (Geogarypidae)

Abstract. Redescriptions of the pseudoscorpion species (Arachnida: Pseudoscorpiones) Olpium lindbergi Beier, 1959 (Olpiidae) and Geogarypus shulovi Beier, 1963 (Geogarypidae) are provided. Additionally, distribution data for ten species belonging to three families, Olpiidae, Menthidae and Geogarypidae, occurring in Lorestan province (western Iran) are provided. Olpium lindbergi is recorded for the first time in Iran.

Pedipalps. Chela distinctly darker in colour than femur and patella; entirely smooth; all setae simple; femur with 2 long tactile setae without enlarged alveoli situated on retrolateral face (Fig. 1c), first seta situated on basal third and second tactile seta situated distal to middle of femur; femur with distinct pedicel, L/W 3.30-3.69; patella with 5 lyrifissures, 4 lyrifissures situated basally, one lyrifissure located ventromedially, L/W 2.53-2.80; chela with distinct pedicel (Fig. 1d-e); chela (with pedicel) L/W 3.77-3.85; chela (without pedicel) L/W 3.50-3.62; hand (with pedicel) L/W 1.59-1.61; movable finger 1.37-1.44 times longer than hand (with pedicel); fixed finger with 8 and movable finger with 4 trichobothria (Fig. 1e); fixed finger with trichobothrium et situated close to tip of finger, it and est situated in the middle of the finger, it located slightly distal to est, ist situated between est and isb and slightly closer to isb, isb situated on retrolateral side of the finger, ib situated basally; movable finger with trichobothrium st situated closer to sb than to t, sb situated in the middle between st and b; fixed finger with 16-18 sensory setae, 5-7 of them situated close to trichobothrium et and fingertip on the distolateral face; fixed finger with 28-30 teeth, 6-7 basal teeth extremely reduced; movable fingers with 22-28 teeth becoming small at the middle between trichobothria t and st; teeth of the movable chelal finger smaller than those of fixed chelal finger; nodus ramosus situated slightly proximal to trichobothrium et in fixed finger and approximately in the middle between tip of finger and trichobothrium t in movable finger (Fig. 1e); primary venom ducts inconspicuous in both fingers. Legs. Not granulate; all setae simple; claws symmetrical, stout and short; arolia simple and much longer than claws, not divided; each coxa I with 4, coxa II with 4-5, coxa III with 4-5 and coxa IV with 6-8 setae (Fig. 1f ) (Beier 1959)], the loss of transverse furrows (Beier 1959), the chelal shape in lateral view (see Beier 1959: Fig. 8), and the structure of the chelal teeth [in the movable chelal finger, cusped teeth are only present in the distal half of the finger (Mahnert 1991)]. The types of O. lindbergi (Beier 1959) are slightly stouter than the specimens found in Iran, e.g. the pedipalpal femur proportion is 3.20-3.30× and the chela (with pedicel) is 3.30-3.50× longer than broad in the types. Loss of some fine granules on the mediodistal face of the chelal hand and the presence of greater number of chelal teeth in the types (34 in the fixed and 30 in the movable chelal fingers) (Beier 1959) are minor small differences between the types and the examined specimens from Iran. These small differences are not sufficient for introducing a different species and can be considered as intraspecific variations within the species. Carapace. Hardly sclerotized; heavily granulated with starlike hispid granulation (Fig. 2a); slightly wider than length or approximately as long as length, L/W 0.94-1.01; front margin elongated distally, snout-like (Fig. 2a), with 14 setae; anterior margin with 4 setae, 2 short setae situated medially and 2 long setae situated sub-medially (Fig. 2b), all setae simple; posterior margin extended laterally, with 6-10 setae; with 2 pairs of well-developed corneate eyes situated away from anterior margin on small protruding mound, anterior eyes slightly larger than posterior eyes, one short seta situated between eyes; anterior furrow present and curved basally; posterior furrow present but indistinct; setae simple; with 6-8 distinct lyrifissures, first pair situated distal to anterior eyes (lost in one female), second pair at same level as posterior eyes, third pair situated closer to anterior furrow than posterior margin and fourth pair situated near posterior margin.
Despite this minor morphometric variation, the only other obvious difference is the position of trichobothrium ib which is located at the same level as est in the female type (see Beier 1963: fig. 7, below), and slightly proximal to est in the male type (see Beier 1963: fig. 7, upper), whereas it is situated distinctly proximal to est in the females from western Iran. Unfortunately, there are no published descriptions about setal shape located on the anterior margin of the carapace and the intensity of granulation on the fixed chelal finger, so these characters cannot be compared in this contribution.
These small differences are not strong evidence for a different species. On the basis of the carapace colouration (bicoloured, different coloured area less contrasted), the absence of wrinkles on the pedipalpal femur and patella, the pedipalpal shape, and the acuminate shape of the galea in females, the newly collected specimens from western Iran are attributed to Geogarypus shulovi which was previously reported from Israel, Iran, Turkey and Turkmenistan (Harvey 2013). Beier (1971) reported this species for the first time from Maku, West Azerbaijan province, north-western Iran (without giving morphometric data), and the presence of G. shulovi in Lorestan province (western Iran) is a new provincial record for the species.
Geogarypus shulovi can be easily separated from G. harveyi Nassirkhani, 2014, the only other species of the genus reported from southern Iran, by the colouration of the carapace (in G. harveyi it is uniformly dark coloured or the posterior half of the carapace uniformly lighter in colour than the anterior half ) and the morphometric data [in G. harveyi, the pedipalpal femur length is 0.69-0.84 mm and the chelal (with pedicel) length is 1.07-1.32 mm (()] (Nassirkhani 2014(Nassirkhani , 2016c.

Discussion
Members of the family Olpiidae are well represented in the province with seven species. The three species Calocheiridius centralis, Cardiolpium bisetosum, and Minniza babylonica are widely spaced from east to west of the province. The occurrence of C. centralis in southwestern parts of Iran has been previously reported by Nassirkhani (2016b). Therefore, this species is widely distributed in western and southern Iran, from Fars province to Lorestan province. Cardiolpium bisetosum may be an endemic species which is distributed from central-west (Nassirkhani 2015) to western Iran. Minniza babylonica and Minniza persica, widely distributed species throughout Iran, can be found in different microhabitats, e.g. under stones, leaf litters and bark pieces (unpublished data).
Cardiolpium asiaticum, Minniza gallagheri and Paramenthus nanus which were previously recorded for Fars province by Nassirkhani (2016aNassirkhani ( , 2016d and Nassirkhani & Vafai Shoushtari (2015), and Geogarypus harveyi which was previously reported from Kerman, Fars and Khouzestan provinces by Nassirkhani (2016c), have been rarely found in Lorestan province. It shows that at least these species are expanded from southwestern to western Iran.