A catalog of the type specimens of various families of Tenebrionoidea (Insecta: Coleoptera) housed at the " Museo de La Plata

. This contribution brings information on the type specimens of Coleoptera Tenebrionoidea deposited at the entomological collection of the Museo de La Plata (MLP). They belong to 29 species or subspecies of the families Aderidae (1 sp.), Anthicidae (11 spp.), Mordellidae (1 sp.), Oedemeridae (1 sp.), Rhipiphoridae (2 spp., 1 subsp.) and Tenebrionidae of the subfamilies Alleculinae (3 spp.), Pimeliinae (5 spp., 2 subspp.) and Tenebrioninae (2 spp.). These types correspond to species described by the specialists C. Berg, C. Bruch, L. Fairmaire, M. Manfrini de Brewer, M. Pic, F. G. Werner and G. Flores. For each species we provide original and valid names, bibliographic reference, category of types, number assigned in the entomological collection of the MLP, sex (if it is indicated) and exact transcription of the labels. There is also additional information on specimens labeled as types but not having this status, because they were compared with types (19 species of Anthicidae, Ciidae, Mordellidae and Tenebrionidae Alleculinae) or corresponding to species seemingly not published.


INTRODUCTION
The superfamily Tenebronoidea includes almost 34,000 species of polyphagous Coleoptera that feed on bark, flowers, fungi and vegetation in general (Slipinksi et al., 2011), or which are associated with carcasses (Aballay et al., 2016).This taxon gathers several families, among them, Tenebrionidae, Meloidae and Anthicidae as the most specious.The type material of Meloidae deposited at the Museo de La Plata has been revised by Campos Soldini et al. (2009) and the types of Tenebrionidae and Perymilopidae, by Cabrera et al. (2010).In this opportunity, we study the type specimens corresponding to the species of Aderidae, Anthicidae, Ciidae, Mordellidae, Oedemeridae and Ripiphoridae, plus some types of Tenebrionidae not included in the catalog of Cabrera et al. (2010) because they were published after this date.
The types of species or subspecies herein treated were described by the specialists Carlos Berg, Carlos Bruch, Leon Fairmaire, Mireya Manfrini de Brewer, Maurice Pic, Floyd Gerald Werner, and Gustavo Flores.The American specialist F. G. Werner contributed with most types (five holotypes and 140 paratypes of nine species of Anthicidae).Berg's collection was originally housed by the National Museum [at present Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" (MACN)] and the University of Buenos Aires, but part of it is currently at the Museo de La Plata (MLP).Moreover, Berg's collection includes several types of species described by Fairmaire (1883).
In order to assist future research we bring additional information on specimens labeled as types but not having this status (e.g. they were collected after the publication of the description or collected in localities not mentioned in the descriptions) and specimens corresponding to species not found in the literature because they are names in scheda or they are nomen nudum (published without a description according to the International Code of Zoological Nomenclature, 1999).

MATERIAL AND METHODS
We examined 290 specimens labeled as types and corresponding to 48 species.For each species we provide the original name and its bibliographic reference, the valid name (between brackets), the geographic distribution (country and provinces or states) of the types examined, the category of types, the sex of the specimens (if it is indicated), the code numbers assigned in the entomological collection of the MLP, the information on the labels, and the color of these labels.The slash after the MLP code indicates if the specimens are holotypes (/1), allotypes (/2), paratypes (/3 and higher numbers) or syntypes (/3 and higher numbers).There is an exact transcription of the labels, in a way that brackets separate different labels and slashes, different lines within each label.Remarks Remarks: Specimen glued on a card along with the abdominal ventrites; genitalia attached in a vial with glycerine; the holotype label is red.Remarks Remarks: The holotype and some paratypes were deposited at the Fundación e Instituto Miguel Lillo, Tucumán, Argentina, and other paratypes are at the Field Museum of Natural History, Chicago, USA.They come from Chaco and Formosa provinces (Argentina) and from Mato Grosso (Brazil).The paratype label is blue.Remarks Remarks: Specimen glued on a card; type label white, with red frame.According to the original publication the type series comes from Argentina, Buenos Aires to Great Chaco.The author did not specify the number of specimens of the type series, however, the vast geographic distribution of the species makes us to conclude that the description was based on more than one specimen.Consequently, the type deposited at the MLP was considered a syntype and was labeled as  Remarks Remarks: Specimens glued on cards; type labels are red.In the original description it is indicated that the two specimens studied for the description of this species, holotype (labeled as typus by the author) and fototype, were deposited at the MLP collection.We interpret that the specimen labeled as type is the holotype, and the fototype is a paratype of sex opposite to the holotype.Both specimens were labeled as such by the authors of this paper.Remarks Remarks: Specimen pinned; type label white, with red frame.The species was described based on two specimens collected by J. Ambrosetti.We were unable Revista de la Sociedad Entomológica Argentina 76 (3-4): [16][17][18][19][20][21][22][23][24][25][26]2017 such by the authors of this paper.

Trigonodera quadrilineata
to find the second specimen in another museum.It is not in the MACN (Bachmann & Flores, 2008)

SPECIMENS LABELLED AS TYPES BUT NOT INCLUDED IN THE ORIGINAL SERIES
We provide information on specimens labeled as metatypes or as homotypes (= "cum typus comparat").Most of them have been collected by Manuel Viana after the publication of the species, and compared with types by him or by the author, Maurice Pic.

Anthicidae
1. Formicilla bruchi Pic, 19041904. Pic.Revista del Museo de La Plata XI, 329.Remarks Remarks: The original publication indicates that the type is from Argentina, Buenos Aires, Bruch col.and was deposited at the Museo de La Plata.This type is probably at the Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" (MACN), which currently holds great part of Bruch's collection.The specimen deposited at the MLP is labeled as metatype and has the following data: Buenos Aires, Tigre, 1938, Viana col.Pic, 19091909. Pic. L'échange 25, 108-109.Remarks Remarks: The original publication indicates that the type is from Brazil, Cuyaba, Pic collection.The specimen at the MLP is labeled as metatype and has the following data: Argentina, Buenos Aires, Isla Martín García, 1936-1937, Viana col. 3. Formicilla strangulata Pic, 1895 1895.Pic. L'échange 11,19.Remarks Remarks: The type comes from Chile, however, Werner (1974)  Remarks Remarks: The type material is from Brazil, without locality information.The two specimens at the MLP are labeled as metatypes and have the following data: Brazil, Santa Catarina, Nova Teutonia, F. Plaumann coll.Pic, 19271927a. Pic. Mélanges Exotico-Entomologiques 48, 23.Remarks Remarks: The type material of this subspecies is from Brazil, without locality information.The two specimens at the MLP are labeled as metatypes and have the following data: Brazil, Santa Catarina, Nova Teutonia, F. Plaumann coll.Pic, 19211921. Pic. Mélanges Exotico-Entomologiques 34, 13-14.Remarks Remarks: According to the original publication the species is from South America, without locality information.The two specimens at the MLP are labeled as metatypes and have the following information: Argentina, Córdoba, II-1939, Viana coll.Pic, 19301930b. Pic. Mélanges Exotico-Entomologiques 55, 18.

Lystronychus ignaciosus
Remarks Remarks: The type material is from Argentina, without locality information.The two specimens at the MLP are labeled as metatypes and have the following information: Brazil, Santa Catarina, Nova Teutonia, F. Plaumann col.There is one syntype from Argentina, Iguazú, X-1928, A. Breyer col., at the Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" (MACN), Buenos Aires (Bachmann & Flores, 2008).

SPECIMENS LABELLED AS TYPES OF SPECIES NOT FOUND IN THE BIBLIOGRAPHY OR WITH UNAVAILABLE NAMES
Four species names are "in scheda" and were probably not published, since we were not able to find them in the literature.Another name, Xystropus femoratus, was published but is unavailable because it was not accompanied by a description.According to Spiessberger et al. (2017) is a nomen nudum.

Mordella patruelis Berg.
Two syntypes from Uruguay, Montevideo.This species name is mentioned in http://www.coleopteraneotropical.org, however, we did not find any description of it.

Xystropus femoratus Klug.
Homotype (= "cum typus comparat") from Uruguay.This species was cited in a list of Alleculinae by Luederwaldt (1911) and in a paper referred to Coleoptera of forensic importance in Brazil, by Massutti de Almeida et al. (2015).According to Spiessberger et al. (2017) it is unavailable, because there is no published description of this species.

10. Ischyropalpus eryngii Werner, 1965 1965
Specimens glued on cards; paratype labels are blue.The holotype was deposited in the National Museum of Natural History, Washington D.C., USA; other paratypes are at the Museu Emilio Goeldi, Belén, Brazil.In the original publication the two paratypes at the MLP are not mentioned.These paratypes were sent by Werner to the Museo de La Plata after the description of the species.

Tenebrioninae 28. Emmallodera rugosa Silvestro and Flores, 2012
. Specimen glued on card.Paratype label is red.The holotype, allotype and some paratypes (all from Catamarca) are deposited in the Instituto Argentino de Investigaciones de las Zonas Áridas, Mendoza, Argentina, and other paratypes are at Fundación e Instituto Miguel Lillo, San Miguel de Tucumán, Argentina; Museo Argentino de Ciencias Naturales "Bernardino Rivadavia", Buenos Aires, Argentina; Field Museum of Natural History, Chicago, USA, and Natural History Museum, Basel, Switzerland.n. sp./ Det.G. Flores &/ J. Pizarro-A.2011].Remarks Remarks: Specimens pinned; paratype labels are red.The holotype, allotype and some paratypes (from Coquimbo region) are deposited in the Museo Nacional de Historia Natural, Santiago, Chile, and other paratypes are at Universidad de Concepción, Concepción, Chile; Laboratorio de Entomología Ecológica, Universidad de La Serena, Chile; Instituto Argentino de Investigaciones de las Zonas Áridas, Mendoza, Argentina; Field Museum of Natural History, Chicago, USA, and Natural History Museum, Basel, Switzerland.
remarked that this species is introduced and probably not established in this country.Formicilla strangulata (=Acanthinus strangulates) is currently distributed in northern Argentina and Uruguay.The specimens at the MLP are labeled as metatypes and Remarks Remarks: Specimen pinned; paratype label is red.The holotype, allotype and some paratypes (from Chubut) are deposited in the Instituto Argentino de Investigaciones de las Zonas ridas, Mendoza,