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8http://zoobank.org/urn:lsid:zoobank.org:pub:E5021D95-8238-4F33-A9F6-0BBE908A2D20
DOI: 10.24412/2226-0773-2024-13-1-21-38
EDN: VDXIRF
Taxonomic notes on longhorned beetles with the descriptions of several new taxa (Coleoptera, Cerambycidae)
M.A. Lazarev
Free Economic Society of Russia, Department of Scientifics Conferences and
All-Russian Projects
Tverskaya str., 22a, Moscow, 125009, Russia
e-mail: humanityspace@gmail.com, cerambycidae@bk.ru
Key words: Coleoptera, Cerambycidae, taxonomy, zoogeography, new genus, new subgenus, new synonyms, new rank, another subfamily added.
Abstract: Many newly published taxonomy modifications are discussed. Several taxonomy news is proposed.
Introduction.
Many taxonomic publications on Cerambycidae are regularly appear each year without special comments by colleagues. I propose here alternative positions on several questionable cases. Several new synonyms cannot be accepted. Several new names were in fact synonyms. Certain taxonomy acts were not acceptable. Rank of six species names was downgraded to subspecies. A new genus and a new subgenus were proposed, as well as many new synonyms. One old subfamily name was restored, as well as one subspecies name. A wrong geographical record was corrected, as well as some other wrongly published data. Rank of one old name was restored.
1. A new unacceptable tribal system of Lepturinae was proposed by Zamoroka (2022b): Cariliini (Carilia, Acmaeops, Gaurotes, Paragaurotes, Dinoptera, Gnathacmaeops, Cortodera); Pidoniini (Pidonia, Fallacea), Evodiini (Evodinus, Brachyta). Lepturini includes 6 Palaearctic genera (Anoplodera, Nivellia, Leptura, Anastrangalia, Grammoptera, Strangalia); Stenocorini - 2 (Stenocorus, Anisorus), Rhamnusiini - 3 (Rhamnusium, Akimerus, Enoploderes), Rhagiini - 2 (Pachyta, Rhagium).
Cortodera cannot be in one tribe with all genera around Dinoptera; Rhamnusium, Akimerus, and Enoploderes are totally different on larval and imaginal characters, so until better decisions it is necessary conserve traditional system of Palaearctic Lepturinae with 7 tribes: Xylosteini, Encyclopini, Oxymirini, Enoploderini, Rhamnusiini, Rhagiini, Lepturini.
2. Evodinellus borealis (Gyllenhal, 1827) = Pidonia petrovi Danilevsky, 2023a, syn. nov. on the base of the type series study.
3. Megarhagium Reitter, 1913 and Hagrium Villiers, 1978 must be accepted as valid genera names as in Villier (1978).
4. Very different genera Brachysomida Casey, 1913 and Pseudogaurotina Plavilstshikov, 1958 were inadequately published by Zamoroka (2022b) as subgenera of one genus, and impossible combination was proposed: “Brachysomida (Pseudogaurotina) excellens”.
5. According to Zamoroka (2022b), “Evodinus borealis does not belong to the separate genus Evodinellus Winkler, 1929” or Evodinus LeConte, 1850 = Evodinellus Plavilstshikov, 1915. But American Evodinus differs from Evodinellus borealis (Gyllenhal, 1827) by the position of antennal insertions, which are situated in Evodinellus borealis in front of anterior eye margins, while in Evodinus antennal insertions disposed behind the line connecting anterior eye margins.
6. The names of very different genera: Acmaeops LeConte, 1850 and Euracmaeops Danilevsky, 2014 were wrongly published by Zamoroka (2022b) as synonyms.
7. Wrong synonyms were accepted by Zamoroka (2022a):
“Cortodera flavimana (Waltl, 1838) = C. moldovana Danilevsky, 1995”. In fact, C. moldovana has no connection with C. flavimana, but close to C. tibialis (Marseul, 1876), and especially to C. tibialis rossica Danilevsky, 2001b. No evidences of the presence of С. flavimana and C. moldovana in Ukraine exist.
8. Cornumutila quadrivittata (Gebler, 1830) was wrongly recorded for Ukraine by Zamoroka (2018, 2022a). C. quadrivittata is widely distributed in Siberia (Lazarev, 2009) and does not penetrate to Europe. A single record for Moscow Region was very doubtful. Only C. lineata (Letzner, 1844) is distributed in Europe.
9. Pedostrangalia revestita corsica Vartanis, 2024b, stat. nov. was described from France (Corsica) as a species on the bases of red anterior legs in combination with black other legs. No other features are observed that distinguish the new taxon from P. r. revestita.
10. The reality of two different subgenera of Stictoleptura [Paracorymbia sensu Miroshnikov, 2021, 2016] subgen. Batesiata Miroshnikov, 1998 and S. subgen. Pyrrholeptura Lazarev, 2016 was not accepted by Miroshnikov (2016, 2021). I insist on three fundamental differences: Pyrrholeptura is characterized by red elytral color, shallow male abdominal emargination and dense elytral punctation. The difference in color between two subgenera can't be denied, but Miroshnikov shows several examples of different taxa where elytral color is not important. Such reasoning is irrelevant. He shows male abdominal emargination in both species, but the shape of those structures in his photos is distinctly different, as well as elytral punctation in his photos of “P. (Batesiata) tesserula” and “P. (B.) pyrrha”. So, Stictoleptura (Batesiata Miroshnikov, 1998) and S. (Pyrrholeptura Lazarev, 2016) are good valid names.
11. Stictoleptura (Maculileptura Danilevsky, 2014) was not accepted by Miroshnikov (2021) who did not see the differences of this taxon from Paracorymbia (s. str.) Miroshnikov, 1998. But that valid name was established by Danilevsky (2014) instead of Paracorymbia (s. str.) «group maculicornis» Miroshnikov (1998). All characters of the group were published by Miroshnikov (1998: 594): Last abdominal male sternite with shallow, narrow, short, but well distinct impression, slightly emarginated apically; hind male tibiae not curved, with two apical spines; elytra yellowish, monochrome or with dark apex and lateral margin; antennal joints with light bases or with light rings.
12. Several new synonyms are proposed: Xylotrechus (Xylotrechus Chevrolat, 1860 = Fulvotrechus Zamoroka, 2021 = Hieroglyphotrechus Zamoroka, 2021 = Igneotrechus Zamoroka, 2021 = syn. nn.), Xylotrechus Chevrolat, 1860 = Spinotrechus Zamoroka, 2021, syn. nov.
In general Zamoroka’s publications were often based on molecular data, but strong morphological argument were usually ignored. For example, such cases as one tribe for so different genera as Rhamnusium, Akimerus and Enoploderes, as well as joining of Rutpela maculata (Poda von Neuhaus, 1761) and Stenurella nigra (Linnaeus, 1758) inside one genus show complete absurdity of his method.
13. Wrong synonyms proposed by Zamoroka (2021): Xylotrechus = Rusticoclytus must be rejected, and valid name Xylotrechus (Rusticoclytus Vives, 1977) generally accepted (Vives, 1977, 2000; Villiers, 1978, 1979; Demelt, 1982; Bílý & Mehl, 1989; Marquet, 2001, 2015; Pesarini & Sabbadini, 2007; Sama, 2008; Löbl & Smetana, 2010; Danilevsky, 2012, 2020; Alekseev & Maryutin, 2019; Stolbov et al., 2019; Trócoli, 2019; Gradinarov & Sivilov, 2020; Sakalian et al., 2020 and others) must be preserved.
14. Xyloclytus was wrongly upgraded to genus rank by Zamoroka (2021). Valid subgenus name Xylotrechus (Xyloclytus Reitter, 1913) must be preserved.
15. Teratoclytus D.W. Zaitzev, 1937 cannot be moved to Anaglyptini, as it was proposed by Zamoroka (2021) and must be returned to Clytini (elytral bases without tubercles).
16. Humeromaculatus Özdikmen, 2011: 537 (type species Cerambyx figuratus Scopoli, 1763) was introduced as a subgenus of Chlorophorus. It was upgraded to genus level by Zamoroka (2021) without sufficient reasons. Sparganophorus Zamoroka, 2021 (type species Clytus diadema Motschulsky, 1854) was described for a single species, which was placed by Özdikmen (2022) in Ch. (Humeromaculatus), so Ch. (Humeromaculatus Özdikmen, 2011 = Sparganophorus Zamoroka, 2021, syn. nov.
17. Perderomaculatus Özdikmen, 2011: 537 (type species Cerambyx sartor Müller, 1766) was upgraded by Zanoroka (2021) to genus level without sufficient reasons, the valid name must be Chlorophorus (Perderomaculatus).
18. Chlorophorus (Viridiphorus Zamoroka, 2021, type species: Callidium herbstii Brahm, 1790) = Chlorophorus (Brevenotatus Özdikmen, 2022, type species: Clytus distinguendus Perroud, 1855), syn. nov. as Ch. herbstii was included in Ch. (Brevenotatus).
19. New synonyms must be accepted:
Stenurella (Priscostenurella Özdikmen, 2013) = Rutpela (Eduardvivesia Zamoroka, Trócoli, Shparyk & Semaniuk, 2022) = Rutpela (Nigromacularia Zamoroka, Trócoli, Shparyk & Semaniuk, 2022), syn. nn.
20. Ropalopus hungaricus olympicus Vartanis, 2024c was described as R. insubricus olympicus Vartanis, 2024c from Greece (Olympos Mt., Pieria prov., 700-1000 m), and R. hungaricus creticus Vartanis, 2024c was described as a species from Crete.
21. Chlorophorus cannot be separated from Clytini, so Clytini Mulsant, 1839 = Chlorophorini Zamoroka, 2021, syn. nov.
22. Three new combinations were incorrectly accepted by Zamoroka et. al. (2022): Rutpela (Eduardvivesia) vaucheri (Bedel, 1900), R. (Nigrostenurella) nigra (Linnaeus, 1758), R. (Nigromacularia) septempunctata (Fabricius 1793). All three taxa must be left in Stenurella as: S. (Priscostenurella) vaucheri, S. (P.) septempunctata and S. (Nigrostenurella) nigra.
Several species were placed by Zamoroka et. al. (2022) in wrong subgenera: Stenurella (Priscostenurella) jaegeri (Hummel, 1825), S. (P.) novercalis (Reitter, 1901), S. (s. str.) hybridula (Reitter, 1902), S. (s. str.) approximans (Rosenhauer, 1856). Must be: S. (Stenurelloides) jaegeri, S. (S.) novercalis, S. (Iberostenurella) hybridula, S. (Crassostenurella) approximans.
23. Oxypleurus nodieri Mulsant, 1839 was placed by Zamoroka (2022a) in Atimiini without sufficient reasons. The species belongs to Saphanini as it is generally accepted.
24. Wrong synonyms were accepted by Zamoroka (2022a):
“Tetropium fuscum (Fabricius, 1787) = T. tauricum Shapovalov, 2007”. The holotype of T. tauricum strongly differs from many hundreds of known T. fuscum. Up to now only very peculiar holotype of T. tauricum is known. So, rather probably it was just a teratic specimen of local species - T. castaneum (Linnaeus, 1758). According to Plavilstshikov (1940), T. fuscum was not collected in Crimea. T. castaneum only was recorded for Crimean fauna by Zagaikevich (1991: 153). In fact T. fuscum is absent in Crimea. It was only recorded by Bartenev (2009) with question mark on the base of WEB European Cerambycidae list of 2000 by M.L. Danilevsky.
25. Anoplistes balcanicus Sláma, 2010 described from Bulgaria was wrongly published as a subspecies of A. halodendri (Pallas, 1773) by Danilevsky (2020) on the bases of the records by Muraj (1960) of Purpuricenus ephippium for Albania. Asias ephippium was also recorded for Bulgaria by Angelov (1995) and for Rumania by Panin, Săvulescu (1961). But in fact, Anoplistes balcanicus Sláma, 2010 does not similar to A. halodendri because of short antenna and legs, peculiar elytral design. So, Anoplistes balcanicus Sláma, 2010 must be accepted as a valid species name, as it was originally introduced. The presence of Anoplistes halodendri in Balcan area rests rather doubtful.
26. Plagionotus detritus caucasicola Plavilstshikov, 1940 = P. d. grecus Vartanis, 2023, syn. nov.
Plagionotus detritus caucasicola Plavilstshikov, 1940 was described with two taxonomy rank in the original publication: as “m.” [morpha] and as “форма” [forma]. The later makes the name available, and it was generally accepted (Özdikmen & Turgut, 2009; Löbl & Smetana, 2010; Danilevsky, 2010; Özdikmen, 2014; Vitali, 2016; Rapuzzi & Sama, 2018; Lazarev, 2019; Danilevsky, 2020; Vartanis, 2023 and others).
Vartanis (2023) accepted P. d. caucasicola Plavilstshikov, 1940 as a valid name, but proposed no distinguishing characters. In fact, the Caucasian populations are just identic to Greece populations. Populations from Peloponnesus were described by Vartanis (2023) and I’ve got good series from Northern Greece (Thessaly): 7 males, 8 females from Ossa Mt. and 1 male, 2 females from near Kalabaka (Meteora). Greece specimens are also rather pale as specimens from Caucasian populations, have same wide yellow pronotum anteriorly, wide yellow elytral stripes, and posterior elytral stripes are partly or totally conjugated.
Acronyms of collections:
MD - collection of M. Danilevsky (Moscow)
ML - collection of M. Lazarev (Moscow)
SM - collection of S. Murzin (Moscow)
ZMM - collection of Zoological Museum of the Moscow State University
Plagionotus detritus caucasicola Plavilstshikov, 1940
Figs. 1-3
Type locality. Russia, Republic of Adygea, Maykop environs (on the bases of lectotype designation).
Description. Dark areas of pronotum and elytra rather light-brown; elytra with widened transverse yellow stripes behind middle, dark stripes in between very narrow; anterior pronotal yellow area very wide, often joined with central transverse pronotal stripe; narrow yellow transverse stripe behind elytral bases more or less reduced or totally absent; body length: 14.3-17.5 mm.
P. d. detritus (Linnaeus, 1758) is characterized by dark-brown ground color of elytra and pronotum; light central pronotal stripe usually well developed; yellow elytral stripes usually very narrow; body length: 10-19 mm (Plavilstshikov, 1940).
Material. Lectotype (Figs 1-2) designated by Danilevsky (2009), published by Lazarev (2019), male (length: 14.0 mm; width: 4.2 mm) with 4 labels: 1) “Cauc. occ. bor. / Maikop / 25.V.[1]928”; 2) “ex coll. Shaposhnikov”; 3) [red] “LECTOTYPUS / Plagionotus detritus / forma CAUCASICOLA / Plavilstshikov, 1940 / M. Danilevsky des., 2008”; 4) [pink] “Зоомузей МГУ (Москва, РОССИЯ) / № ZMMU Col 00180 / Zool. Mus. Mosq. Univ. / (Mosquae, ROSSIA) / ex coll. N. N. Plavilstshikov” - ZMM; Paralectotype (Fig. 3): 1 female, Maykop, 5.6.1935 - ZMM.
Additional materials. 1 male, Krasnodar Reg., L’vovskaya, 16.6.1966 - ZMM; 1 female, Ekaterinodar, 9.7.1914, Lyutkovsky - ZMM; 1 male, 1 female, Stavropol, V. Lutshnik - ZMM; 1 male, Armenia, Dilizhan, 5000’, 26.7.1934, N. Plavilstshikov - ZMM; 1 female, Transcauc., Kars - ZMM; 1 male, Krasnodar, Novoprokhladnoe, 13.6.1956-1959 - MD; 1 female, Sochi, Lazarevskoe, 4.8.1983, A. Koval - MD; 1 male, Krasnodar, Ubinskoe, 10.5.1976, M. Kravchenko - MD; 1 male, Krasnodar, Ubinskoe, 2.5.1991, A. Abramov - MD; 1 male, Krasnodar, Ubinskoe, 5.8.1976, Belov - ML; 1 male, Krasnodar, Novoprokhladnoe, 25.5.1959 - MD;. 3 males (Fig. 4), 4 females (Fig. 5), Ossa Mt. (East), VII.2001, P. Tauzin - ML; 4 males, 4 females, with the same label - SM; 1 male, 2 females, 1-2 km N Kalabaka, Meteora, VI, 1981, M. Sláma - ML.
Distribution. Russia, North Caucasus, Georgia, Armenia, Azerbaijan, Iran, Turkey; Greece (Thessaly, Peloponnesus); the records for Syria (Plavilstshikov, 1940; Danilevsky, 2020; Vartanis, 2023) were most probably connected with P. detritus africaeseptentrionalis Tippmann, 1952.
27. Dorcadion fulvum erythropterum Fischer von Waldheim, 1823 = Dorcadion fulvum opillicum Zamoroka, 2019, syn. nov. Big available series of the species (including series from “Opillya” - geographic region of the Podolian Upland in Lvov Oblast, Ivano-Frankovsk Oblast and Ternopol Oblast in western Ukraine) show a great degree of geographical variability masking local forms.
28. Dorcadion fulvum heracles Vartanis, 2024a, stat. nov. was described from Greece (Olympus Mt., Pieria prov.) as a species on the bases of black first antennal joint and black anterior legs. No other features are observed that distinguish the new taxon from D. f. fulvum (Scopoli, 1763).
29. The new wrong synonyms were published by Zamoroka (2022a): without analises of corresponding materials and with false statement: “ranges of some of them completely overlap”. “Dorcadion cinerarium cinerarium (Fabricius, 1787) = D. c. macropoides Plavilstshikov, 1932 = D. c. zubovi Lazarev, 2011”, “D. c. panticapaeum Plavilstshikov, 1951 = D. c. bartenevi Lazarev, 2011 = D. c. skrylniki Lazarev, 2011 = D. c. azovense Lazarev, 2011 = D. c. gorodinskii Danilevsky, 1996 = D. c. demidovi Danilevsky, 2013 = D. c. mosyakini Danilevsky, 2021“; “Dorcadion equestre (Laxmann, 1770) = D. e. vadimi Danilevsky, 2021”; “Dorcadion holosericeum Krynicki, 1832 = D. h. ustinovi Danilevsky, 2021”. Most probably Zamoroka did not know such specimens, so, valid names must be preserved: D. c. macropoides Plavilstshikov, 1932; D. c. zubovi Lazarev, 2011; D. c. bartenevi Lazarev, 2011; D. c. skrylniki Lazarev, 2011; D. c. azovense Lazarev, 2011; D. c. gorodinskii Danilevsky, 1996; D. c. demidovi Danilevsky, 2013; D. c. mosyakini Danilevsky, 2021; D. e. vadimi Danilevsky, 2021; D. h. ustinovi Danilevsky, 2021.
30. The wrong synonyms Falsomesosella truncatipennis Pic, 1944 = F. taibaishana Lazarev, 2021 published by Lin, Weigel & Ge (2021), can’t be accepted, as holotype of F. truncatipennis (depicted by Lin et al., 2021) from Zhejiang (see Lin, 2015) is distinctly wider with more elongated prothorax, besides its type area is strongly distant. So, F. taibaishana Lazarev, 2021 is a valid name.
31. Quasimesosella ussuriensis was recorded by Danilevsky (2023b: 334) for the south of Khabarovsk Region (Listvyanaya River), but according to personal communication by N. Anisimov (November, 2023), that record must be connected with Listvenichnaya River from Malyi Khingan in Jewish Autonomous Republic. The record of that species for Duchin must be connected with Dichun River south-eastwards Radde (Jewish Autonomous Republic).
32. Paratetrops gen. nov.
Type species. Tetrops warnckei Holzschuh, 1977.
Description. Body densely covered with long thick black erect setae; antennae extremely sick, with apical joints about as long as wide; pronotum with smooth central stripe; elytra with rough big punctation. A single species distributed in south Turkey is known - Paratetrops warnckei (Holzschuh, 1977), new rank.
Etymology. close to Tetrops. Gender masculine.
33. Tetrops peterkai Scořepa, 2020 is downgraded to subspecies rank: Tetrops praeustus peterkai Scořepa, 2020, stat. nov.; type locality: Czech Republic, Moravia, Horní Pole environs.
According to the original description the species is distributed in Czech Republic, Slovakia, Austria, Germany.
34. Tetrops praetermitus Sláma, 2020a is downgraded to subspecies rank: Tetrops praeustus praetermitus Sláma, 2020, stat. nov.; type locality. Bohemia, Lásenice. According to Sláma (2020b) the taxon is distributed in South Bohemia only.
35. Phytoecia (Musaria) rubropunctata (Goeze 1777) was wrongly recorded for Ukraine by Zamoroka (2022a). It is a West European species absent in Ukraine; it does not penetrate east of Germany (Bense, 1995; Sama, 2003), but was recently recorded for Spain, France and Italy only (Löbl & Smetana, 2010; Danilevsky, 2020). Old wrong published records could be based on specimens of Ph. (M.) argus (Frölich, 1793) or Ph. (M.) faldermanni (Faldermann, 1837).
36. A new subgenus Phytoecia (Danilevskia subgen. nov., type species: Saperda molybdaena Dalman, 1817) is proposed for 4 species: Ph. (Danilevskia) molybdaena (Dalman, 1817), Ph. (Danilevskia) uncinata (W. Redtenbacher, 1842), Ph. (Danilevskia) tenuilinea Fairmaire, 1876 and Ph. (Danilevskia) badenkoi Danilevsky, 1988. The new taxon is characterized by the absence of the dense solid elytral scaly cover consisting of small scales or very short setae. Such cover is typical for Ph. (Opsilia Mulsant, 1862). Besides all Ph. (Danilevskia) has unicuspid mandibulae and eyes with joined dorsal and ventral lobes. New subgenus name is feminine. The name is dedicated to Mikhail Leontievich Danilevsky - a specialist on Palaearctic Cerambycidae and my constant colleague.
37. Up to now Palaearctic Cerambycidae includes 8 subfamilies (Parandrinae, Prioninae, Lepturinae, Necydalinae, Spondylidinae, Apatophyseinae, Cerambycinae, Lamiinae). Now another one must be accepted: Agapanthiinae Mulsant, 1839 (type genus Agapanthia Audinet-Serville, 1835, monobasic), which was traditionally included in Lamiinae.
Agapanthiinae is characterized by moderately or small body size; usually strongly elongated; parallel sided or with sides slightly diverging posteriorly; antennae usually long, often much longer than body, usually 12-segmented (in Pseudocalamobius -
11-segmented); frons usually sloping backwards; prothorax always without lateral spines or tubercles; legs short; anterior coxae spherical; claws simple, divergent, without tooth-like appendages or denticles; metepisternae very narrow, parallel sided. Larvae with cylindrical “C”-like curved body, without legs; head slightly elongated, strongly prominent; abdominal ventral ampullae partly reduced.
Agapanthiinae differ from Lamiinae by many larval characters: “C”-like curved body, slightly elongated head, rounded laterally, ventral ampullae partly or totally reduced.
38. Agapanthia kindermanni Pic, 1905 must be returned to the original subspecies rank as Agapanthia dahli kindermanni Pic, 1905.
39. Agapanthia lateralis Ganglbauer, 1884 is downgraded to subspecies rank: Agapanthia dahli lateralis Ganglbauer, 1884, stat. nov.
40. Agapanthia mutinensium Sama & Rapuzzi, 2010 is downgraded to subspecies rank: Agapanthia dahli mutinensium Sama & Rapuzzi, 2010, stat. nov.
41. Agapanthia pustulifera Pic, 1905 is downgraded to subspecies rank: Agapanthia dahli pustulifera Pic, 1905, stat. nov.
42. Agapanthia salviae Holzschuh, 1975 is downgraded to subspecies rank: Agapanthia dahli salviae Holzschuh, 1975, stat. nov.
43. Agapanthia schurmanni Sama, 1979 is downgraded to subspecies rank: Agapanthia dahli schurmanni Sama, 1979, stat. nov.
44. Agapanthia subsimplicicornis Sama & Rapuzzi, 2010 is downgraded to subspecies rank: Agapanthia dahli subsimplicicornis Sama & Rapuzzi, 2010, stat. nov.
45. Zaitzev D.W. (original spelling - Zaitzev 1931, 1937) was wrongly published several times (Löbl & Smetana, 2010; Danilevsky, 2020) as Zaitzev D.A. The original spelling was unacceptably changed by Zamoroka (2021) to “Zajciw”.
Acknowledgements. I am very grateful to M. Danilevsky (A.N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia), A. Gusakov (Zoological Museum of Moscow State University, Moscow, Russia) and S. Murzin (Moscow, Russsia) for supplying me with material for study. My special thanks to M. Danilevsky for permanent consultations.
Figs 1-5. Plagionotus detritus caucasicola Plavilstshikov, 1940:
1. Lectotype, male; 2. Lectotype labels; 3. Paralectotype, female, Maykop, 5.6.1935; 4. Male, Ossa Mt. (East), VII.2001, P. Tauzin;
5. Female with the same label.
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Received: 20.12.2023
Accepted: 29.01.2024
