Fancy red carp accumulated ingested astaxanthin in the integument. Zeaxanthin was converted to astaxanthin de novo. Lutein was converted not to astaxanthin, but to doradexanthin.
Nippon Suisan Gakkaishi is an official journal of the Japanese Society of Fisheries Science written in Japanese only. The society publishes an English journal, Fisheries Science, as well, which accepts submissions from non-members of the society over the world. Detailed information is available in http://jsfs.jp/en/journals/fisheries-science.