Impacts of climate change on dragonflies and damselflies in West and Central Asia

To project the impact of climate change on dragonfly and damselfly diversity in West and Central Asia.


| INTRODUC TI ON
Global biodiversity is in decline largely because of anthropogenic forces and climate change. However, not all ecosystems are changing at the same rate: freshwater biodiversity is declining faster than both marine and terrestrial biodiversity (McRae et al., 2017). The causes of such loss to freshwater bodies are disruptive pressures like flow regulation, habitat degradation, pollution, species invasion, species overexploitation and climate change (Dudgeon, 2019;Grill et al., 2019;IPBES, 2019); and the severities of these effectors are expected to increase with projected human population growth.
Freshwater covers only <1% of Earth's surface, and amounts to ~0.007 of the total planetary water supply (Reid et al., 2019) but hosts almost 9.5% of all described animal species (Balian et al., 2008), including an estimated 70 mammal species, 5700 dragonflies (Balian et al., 2008) and 18,495 fishes (Fricke et al., 2023).  (Cohen et al., 2016;Pelicice et al., 2017) and temperate (Freyhof & Brooks, 2017) scales. According to the current IUCN Red List, approximately 30% of freshwater species are threatened with extinction; therefore, it is imperative that researchers monitor these ecosystems' population and distribution changes, including through the use of barometric indicator species.
Odonata, including dragonflies (Anisoptera) and damselflies (Zygoptera), can serve as effective model organisms for freshwater biodiversity. This is possible because of their: (a) ease of identification, (b) high diversity, (c) rapid development, (d) complex life histories and (e) essential roles within food webs (Collins & McIntyre, 2015;Corbet, 1999;Simaika & Samways, 2011). The order is also a viable climate change indicator because some of its species' geographical distributions (Hassall & Thompson, 2010;Hickling et al., 2005), development rates, body sizes, colours (De Block & Stoks, 2003;Hassall & Thompson, 2008) and phenologies (Hassall et al., 2007) change with increasing temperature. Furthermore, their range of adaptations, which have allowed them to inhabit temperate to subarctic areas ever since their Lower Permian, tropical origins, will likely enable them to outlast future climate change. Despite this, local populations are still susceptible to extinction by climate change because as they relocate their niche space will decrease with higher latitude and elevation (Hassall & Thompson, 2008). Odonata also can indicate freshwater quality and supply because they need waters that are well connected, lowly polluted and resource rich (Angelibert & Giani, 2003;Clark & Samways, 1996;Hassall & Thompson, 2008;McCauley, 2006;Pither & Taylor, 1998;Raebel et al., 2012). The order also has the advantage that its taxonomy and distribution are well known compared to most other freshwater groups, especially invertebrates (Bried et al., 2020); over the past 20 years, a good amount of data have been published globally on the increasing effects of climate change on odonate distribution patterns (Bush, Nipperess, Duursma, et al., 2014;, Hickling et al., 2005, faunal compositions (Flenner & Sahlén, 2008) and phenologies (Dingemanse & Kalkman, 2008;Hassall et al., 2007).
Regarding Central Asia's projected climate change, temperature may increase by 0.23-0.4°C per decade, depending upon the climate scenario modelled (He et al., 2021); the results of precipitation models vary but do not project strong changes (Lioubimtseva & Cole, 2006).
From the expected changes in these regions by century's end, local freshwater diversities and distributions are at a high risk of disruption, which means conservation measures are urgently needed.
Since Odonata inhabit these regions, they can indicate the quality of these freshwaters.
To project the geographic distribution of a certain species in time, whether present or future, species distribution modelling (SDM) is often used. SDMs predict areas that possess suitable habitat for a species by correlating their occurrences with climate variables and topographic information (Beilinson, 2016;Elith & Leathwick, 2009).
These environmental variables characterize places where a species has been recorded and then the SDM delimits similar areas suitable for habitation. SDMs are, however, limited in that they rarely account for biotic interactions, such as inter-species competition, meaning their projections for given taxa could be uninhabitable (Finch et al., 2006). In the past, distributions of freshwater taxa, including Odonata, have been successfully projected using SDMs, and often resulted in climate change-induced range shifts (Bush, Nipperess, Duursma, et al., 2014;Domisch et al., 2011;Maes et al., 2010).
Odonata are well suited for SDMs because their occurrence data are relatively unbiased compared to other invertebrates since their charisma and visual identifiability promote large data sets, which are suitable for the framework used here (Collins & McIntyre, 2015).
To gain insight into how dragonfly and damselfly biodiversity may be impacted by climate change, we modelled the expected distribution of 159 species occurring in West and Central Asia, an area encompassing 24 countries and spanning 8 million square kilometres. We did so for three climate scenarios (SSP1-2.6, SSP3-7.0 and SSP5-8.5) for two different time periods (2050-2070 and 2080-2100). These scenarios give us insights into what might happen in a best-case scenario where the targets of the Paris Agreement are met (SSP1-2.6), a worst-case scenario where no climate action is taken (SSP5-8.5) and an in-between scenario (SSP3-7.0) which is currently believed to be the most likely outcome. Based on these models and the diversity maps derived thereof, we test four hypotheses: 1. Climate change in West and Central Asia will favour habitat generalists that can use either lotic or lentic habitats over habitat specialists that require lotic habitats. Increasing drought is expected to result in an increasing number of brooks and rivers to become intermittent during parts of the year, rendering them unsuitable for many lotic species but still suitable for reproduction by habitat generalists capable of successful reproduction in standings waters.
2. Climate change in West and Central Asia will result in an increase in species with an Oriental and Afrotropical distribution and a regional decline in species with a Palaearctic distribution. Increasing temperatures are likely to favour the species whose distribution is centred on the largely tropical Oriental and Afrotropical regions while negatively impacting species adapted to more temperate climates and dependent on more boreal habitats such as bogs. We additionally expect that these changes in ranges will coincide with changes in the connectivity (patch size and patch area) of suitable habitat for species with different origins.
3. Climate change in West and Central Asia will lead to an increase in dragonfly and damselfly biodiversity gaps (areas where dragonflies and damselflies are largely absent). In large areas of West and Central Asia, permanent freshwater habitats are scarce and diversity of dragonflies and damselflies is largely dependent on regular colonization. The increase in temperature due to climate change is expected to increase the extent of these low-diversity areas.
4. Endemic species will on average be more severely negatively impacted by climate change. Many of the species endemic to West and Central Asia are confined to mountain ranges, often occurring at higher elevations. To adapt to the increasing temperatures, these species need to shift to higher elevations, which will likely result in a strong decline in suitable area.

| Study area
The study area covered West and Central Asia at the crossroads of the Palearctic, Afrotropical and Oriental realms. The countries included Turkey, Cyprus, Israel, Palestine, Syria, Lebanon, Saudi Arabia, Oman, Yemen, the United Arab Emirates, Georgia, Armenia, Azerbaijan, Iran, Iraq, Tajikistan, Kyrgyzstan, Uzbekistan, Turkmenistan and Afghanistan. Kazakhstan was excluded from this study because it is too large and poorly explored.

| Species data
The taxonomy follows  with the exception that Calopteryx waterstoni is regarded as a subspecies of C. splendens. In total, 172 species of dragonflies and damselflies have been recorded from the study area (Table S1). Of these 172 species, 13 were discarded as they are either only known from vagrants from other parts of Africa or Asia or are only known from one or two localities due to which their distribution could not be modelled (Acisoma panorpoides, The 159 modelled species were, based on expert knowledge and literature (Boudot et al., 2009, Boudot & Kalkman, 2015, divided into two groups, habitat specialists that require lotic habitats (56 species) and habitat generalists that can use either lotic or lentic habitats (103 species). Species were considered lotic species when they are dependent on running water habitats (rivers, brooks, wadis, seepages, etc.) and would go largely or completely extinct when no running water would be available. All other species are considered lentic species, meaning that this group also includes species that regularly reproduce in running water without being dependent on it.
Furthermore, the species were also divided into groups based on their biogeographical origin. This was based on their present distribution or the distribution of their nearest relatives. Thirty-nine species were considered of Afrotropical origin, 16 of Oriental origin and 93 of Palaearctic origin. A further 11 species could not be placed in one of these categories, mostly as they are wide ranging across the tropics of both the African and the Oriental regions.

| Modelling framework
Climate data were obtained in the form of the 19 bioclim variables, along with the minimum, mean and maximum temperatures per month, and mean precipitation per month for the period 1980-2010 from CHELSA (Climatologies at high resolution for the Earth's land surface areas; Karger et al., 2017). We opted to use temperature and precipitation variables because they are known to influence odonate thermoregulation and hydroperiod (Collins & McIntyre, 2015).
Collinearity between variables was assessed using a Pearson correlation coefficient of <0.7, and those that were highly correlated were chosen based on greatest ecological relevance. The final selection included annual mean temperature (BIO1), temperature seasonality (BIO4), annual precipitation (BIO12), precipitation seasonality (BIO15), climatic moisture index, potential evapotranspiration (PET) driest quarter and PET wettest quarter. River and soil data were also used, including total river length (m) and percentage cover of sands, aridsols and entisols. The river data were obtained from HydroSHEDS HydroRIVERS Version 1.0 and the United States Department of Agriculture (USDA) Natural Resources Conservation Service (Linke et al., 2019) and the soils data from the global soil regions map from the USDA Natural Resources Conservation Service Soils (USDA, 2005). The environmental variables were aggregated at a spatial resolution of 0.4 decimal degrees (~45 km) to match the precision of the species collection data. Future climate data were also obtained from CHELSA from CMIP6 climate models. The same variables were collected for three scenarios of shared socioeconomic pathways (SSPs) for the years 2050-2070 and 2080-2100. The three scenarios were as follows: SSP1-2.6, SSP3-7.0 and SSP5-8.5.
The first scenario projects a low greenhouse gas emission level (−9 GT CO2 by 2100 relative to pre-industrial levels) with a global average surface warming of 2°C; SSP3-7.0 projects a moderate emission level (83 GT CO2 by 2100) with a warming of 4°C; and SSP5-8.5 projects a high emission level (126 GT CO2 by 2100) with a warming of 5°C. Each variable was calculated from mean temperature and rainfall data aggregated from five global circulation models: GFDL-ESM4, IPSL-CM6A-LR, MPI-ESM1-2-HR, MRI-ESM2-0 and UKESM1-0-LL. We assumed that by these future years, river and soil presence will remain the same.
For each species, we conducted the species distribution modelling using R statistics (V.4.1.2; R Core Team, 2020). We ran each model using the 'ENMeval' package (v2.0.1; Kass et al., 2021). We modelled the distribution for each species using maximum entropy (MaxEnt; Phillips et al., 2006). We chose to use MaxEnt because it performs well for similar data on flying insects (Aguirre-Gutiérrez et al., 2013) and is robust against overfitting (Aguirre-Gutiérrez et al., 2013;Phillips et al., 2006). Absences were only taken from areas where dragonflies had been sampled previously, known as target group sampling (Mateo et al., 2010). Only linear and quadratic features were used for species with fewer than 100 records and hinged features were added for species with more than 100 records. Model performance was measured using average AUC from spatially independent cross-validation, which has been shown to provide models which transfer better to new conditions (Roberts et al., 2017). We used k-means clustering to assign each occurrence cell and background cell into discrete spatial blocks and obtained values of the average model performance using each spatial block as testing data (Kass et al., 2021). Species with fewer than 20 records were clustered into two spatial blocks, fewer than 50, three blocks and those with more than 50 records into four blocks. The single best model for each species was selected as the model with the lowest Akaike information criterion (AIC). We compared the average AUC value of each model to a one-sided 95% confidence interval of the null distribution of average AUC values within the same model and blocking structure (Raes & ter Steege, 2007), keeping only the species whose average AUC value was higher. Each best model was then projected onto West and Central Asia for the present and the three future scenarios in 2070 and 2100. We converted each mapped habitat suitability into a binary map using the largest threshold that would leave out a maximum of 10% of the occurrence records. We compared the future distribution patterns to the present, generating range change statistics (km 2 ) using the 'biomod2' package (v3.5.1; Thuiller et al., 2021).
We prepared three kinds of maps to visualize present and future (2070 and 2100) distributions of each species group (Palearctic, Afrotropical and Oriental). They showed: one, absolute species count; two, the difference in species gained and lost as an area in absolute count; and three, the proportion of species change by area.
The second map type was made by subtracting a species group from that same group's present data to visualize its change from the present. The third map type, bearing proportional change, was calculated per species group by dividing the difference between species group and that group's present location by that species total count, and then multiplying it by 100; thus, what resulted were percentage changes in a species group per area.

| Statistical analyses
To determine the per cent gain or loss of every species for each future year under all three climate scenarios, we divided the area of the future range size (km 2 ) at full dispersal by the current range size (km 2 ). This provided the amount of territory gained or lost in a percentage of the current distribution per species under full dispersal.
For climate scenario SSP 3-7.0, species were divided into groups of severity of change, either increase or decrease, based on the per cent of territory lost or gained for both 2070 and 2100. The species were placed in five different trend categories: strong decline (predicted range is 60% or less of the original range), decline (predicted range 60%-80% of the original range), stable (predicted range is 80%-120% of original range), increase (predicted range is 120%-140% of original range) and strong increase (predicted range is over 140% of the original range). The percentage of each of these categories was then taken from the whole of each species group type to show the proportion they hold of the total species group. The dispersal dependency, defined as the percentage of its future range not being part of its current range, of every species for each year and climate scenario was also calculated by dividing the territory gained from the future range size at full dispersal divided by 100.
This gives an indication of what extent the survival of a species is dependent on its dispersal capacity. A Fisher's exact test was used to test for differences in trend categories between species of standing or running water (hypothesis 1), species of different biogeographical realms (hypothesis 2) and between endemic and non-endemic species (hypothesis 4).
To further test the spatial reorganization of the Odonata community projected under climate change, we explored how species range change impacts the overall connectivity of suitable habitat patches per species in the year 2080 under ssp370. We used patch size and patch area to separate the different ways in which species were predicted to increase or decrease in range, that is, does range change result in larger patches of suitable habitat and are there more or less patches? We fit two linear models to test two separate null hypoth-  (Figure 2a), but in the future, percentages of species present will change by 10%-30% in the areas stated previously. Palaearctic species, which currently within our study area primarily inhabit Turkey (Figure 2b), will lose the greatest distribution in these areas, as well as in Iran and the central part of Central Asia, but gain territory in eastern Turkey and eastern Central Asia (Figure 2h,n). Oriental species on the other hand gain considerable territory (up to 40% increases) throughout the region (Figure 2j,p) from their current primary inhabitants of the Tigris-Euphrates River system (Figure 2d). Afrotropical species will likely remain in western Saudi Arabia, Yemen and Oman (Figure 2c), however, may increase in prevalence therein; they may also increase in western Turkey, southern Iran and the middle of Central Asia (Figure 2i,o).
Lentic species that live throughout the Middle East, and primarily in Turkey (Figure 2e), will likely lose considerable land in western Turkey, the Levant, Georgia, Azerbaijan and northern Iran (10%-20% losses) (Figure 2k,q). Lotic species today live throughout Turkey, Iran and Afghanistan (Figure 2f). In the future, 10%-30% of their diversity in parts of the Eastern Mediterranean, Azerbaijan and Iran may be lost. They may, however, see an increase within eastern Turkey, Armenia, Georgia and eastern Central Asia (Figure 2l,r). Table 1 presents the counts of species per cent change in the future relative to the present under climate scenario SSP 3-7.0. The full data of species per cent change can be found in Table S2. Table 1 further shows the per cent of species per group total per intensity of change to convey the proportion of species falling, rising or remaining stable in the future. From the table, it is apparent that approximately half of all the species modelled will remain 'Stable' by 2070 (54%) and 2100 (40%). The remaining species of the all-species group decline (total 23% for 2070; total 31% for 2100) or increase (total 23% for 2070; total 29% for 2100) about evenly. Of the species modelled, the Palaearctic are predicted to decline the most, with a total of 61% decreasing by 2100, compared to a total increase of 16%. The 13 Oriental species modelled will populate the region studied with a 'Strong Increase' of 57%. Afrotropical populations are largely expected to remain the same (69% 'Stable' for 2070), except for a total increase of 36% by 2100 and a much smaller decrease (total of 6%). Lentic and lotic species, which include Palaearctic, Afrotropical and Oriental species, show a balanced and stable trend respectively. Lentic populations will most likely rise (total 30% for 2100) and fall (total 37% for 2100) in the region about equally, while lotic species may remain generally 'Stable' (51% for 2100). Endemic species (18 total) are predicted to increase (total of 45% for 2100) in the region, and non-endemic species should distribute evenly (total 27% loss and 32% gain for 2100). Figure 3 show the change in grid cells with 15 species or fewer for the three climate scenarios in the years 2070 and 2100. The table shows a decrease in total dragonfly and damselfly biodiversity gaps in the region from 509 grid cells in the present to on average 421 and 418 grid cells in the years 2070 and 2100 respectively. Thus, according to our data, the prevalence of dragonfly and damselfly biodiversity gaps will decline (−18% for 2100) in the region's future.

Table 2 and
Regarding our four hypotheses, expectations were met but there were also surprises. One, we did not find statistical support for the hypothesis that lotic species distributions will decrease more than lentic species distributions; the distribution of lotic species will likely remain more stable (year 2070 χ 2 = 8.466, p-value = .062; year 2100 χ 2 = 7.821, p-value = .111). In fact, some populations of the latter group decrease more than the former, but that is balanced by a proportional increase in other lentic species. Two, Oriental and Afrotropical distribution will likely increase, while Palaearctic distribution decreases, which is the scenario that we assumed (year 2070 χ 2 = 16.736, p-value = .002; year 2100 χ 2 = 34.45, p-value = .001).
Three, climate change is predicted to lead to a decrease of 18% in dragonfly and damselfly biodiversity gaps, which contradicts our expectation for an increase of low-diversity areas. And four, endemic species distributions will broaden their ranges, rather than be neg-  Figure 4a). However, as both these relationships are non-significant, we cannot reject the null hypothesis that there is no change in the number of patches with range change. In terms of mean patch area, there is a strong positive relationship (β = −0.38, p < .001; Figure 4b). An increase in mean patch area is strongly associated with an increase in range and this does not vary based on the origin of the species. Overall, the lack of increase/decrease in the number of patches and the strong decrease in patch areas for declining species implies that patches become smaller but do not disappear, potentially representing refugia. The model results also support the conclusions observed with Fisher's exact test for aggregate species trends, and Palaearctic species are projected to lose significantly more range than Afrotropical and Oriental species, which were projected to increase in range ( Figure 4c). Table S2 per species group type for years 2050-2070 and 2080-2100 under climate scenario SSP 3-7.0; and per cent of total species per group type.

| DISCUSS ION
The discussion will focus on the SSP 3-7.0 climate scenario in 2070 and 2100, which is at present considered the most likely future socio-economic pathway given current trends (IPCC, 2021). become more fragmented but that existing suitable habitats will shrink, and they will become restricted to smaller refuges. This makes them more susceptible to the impacts of mismanaged freshwater habitats.

| Habitat preferences and trends
We expected that species of running water would on average be more negatively impacted than those of standing water, but our predictions show no significant difference in trend between these groups. There are, however, clear spatial differences in the impact of climate change on the diversity patterns of these groups. The decline in lotic species concentrates on southern Turkey (basically the Taurus mountains), the Levant and the southern part of Iran (basically the Zagros mountains). In lentic species, losses are expected mainly in western Turkey, the Levant and the interior of Iran. There is a surprisingly strong difference in impact on lentic species in the Arabian Peninsula with a large area where a decline is expected in the Levant and northern parts of the Arabian Peninsula and a large area where increase is predicted covering most of the south of the Arabian Peninsula. This seems to be correlated with differences in the climate change impact with the whole region getting warmer but the south in addition also getting wetter and the Levant area getting dryer (Almazroui et al., 2020). We would expect that a finer-scale model, which considers distribution patterns at the spatial scale of freshwater bodies, would elucidate any differences in range expansion/contraction of species adapted to standing and running water.
Our results show that these differences in habitat do not reflect coarse-scale trends, but we may be overestimating the range of some species by aggregating fine-scale freshwater maps (Fournier et al., 2017). The increase in species with a mainly African distribution has already been going on in Europe for the past 30 years (Kalkman et al., 2018;Termaat et al., 2019). In Turkey and Cyprus, an increase in African species is also evident (Sparrow et al., 2021) and

| Faunal composition and dragonfly and damselfly biodiversity gaps
it seems likely that this increase is also already taking place in other parts of our study area. Evidence for an increase in Oriental species already taking place in our area is limited, although data suggest that Trithemis aurora has expanded its range westwards along the Persian Gulf coast of Iran in recent decades (Schneider & Ikemeyer, 2019). It is, however, likely that this lack of evidence of increase in Oriental species is mostly due to the relatively scant data for this part of the region due to which it is difficult to dis- increase or strong increase is dominated by species belonging to the families Coenagrionidae and Libellulidae. This is partly due to these families being the most species rich, but these families are also known for containing a relatively high number of species with large ranges, strong dispersal capacity, resilience to saline conditions and are often dominant in unshaded habitats with standing water both artificial and natural habitats (Suhling et al., 2015), which makes them well adapted to arid environments with temporary waters.
In contrast to expectations, an 18% decrease instead of an increase in dragonfly and damselfly diversity gaps was found (areas with <15 species). This decline in low-diversity areas is caused by areas at higher elevations in Central Asia becoming available to odonates due to the increased temperature and due to the increased precipitation in the Arabian Peninsula outweighing the impact of the higher temperature.

| Impact on endemic species
Our test gave no indication that species endemic to our region are more strongly impacted by climate change than non-endemic

| Comparisons to existing knowledge
Our results agree with previous research that odonate distribution and vulnerability will be impacted by climate change (Basel et al., 2021;Bush, Nipperess, Duursma, et al. 2014;Jaeschke et al., 2013;Pires et al., 2018). In our study, the results clearly show that Palearctic species will largely leave the area or become locally extinct, while Oriental and Afrotropical species will replace them. In past research, it has been observed that Odonata distributions shift poleward or contract based on species type (Jaeschke et al., 2013;Pires et al., 2018;Simaika & Samways, 2015). According to Bush, Nipperess, Duursma, et al. (2014),

| CON CLUDING REMARK S
Our predictions are based on data on dragonflies and damselflies, but it is likely that similar patterns of change can be expected in other freshwater groups such as fishes, mollusc, Ephemeroptera, Plecoptera, Trichoptera, aquatic bugs and even freshwaterdependent plants. There is, however, one key difference, and that is the far more limited dispersal capacity of most of these groups.
Especially fish and molluscs but also in Ephemeroptera, Plecoptera and Trichoptera dispersal capacity is such that waters divided by barriers of a few dozen kilometres will not be colonized. In the case of dragonflies and damselflies, we saw that on average 16 per cent of a species potential area of distribution in the future is dependent on dispersal which gives an indication of the additional threat of climate change to species groups with no or limited dispersal capacity.
In this study, we model the impact of climate change. What we do not model is the added impact of mismanagement of freshwater habitats. Already freshwater habitats are strongly under pressure resulting in the decline in freshwater diversity throughout the region and it seems likely that climate change will not only result in a change in the climate but also an increase in human pressure on the remaining freshwater sources both on a local scale such as farmers increasing the extraction of freshwater from brooks and on a national scale where countries try to regulate river systems to their own advantage. This most likely means that the decline in some species is underestimated. An example of such a species is Onychogomphus macrodon, a species restricted to larger river systems in the Levant. Water pollution, gravel mining and changes in river regime due to the construction of dams have already led to a severe decline in this species. Although the model predicts the climatologically suitable area to show an increase, it is far more likely that the past decline in combination with the expected decline due to a further increase in construction of dams in larger rivers and the intake of water will result in this species experiencing further declines maybe to the point of going extinct .
The results show that climate change will become the main driving force behind changes in dragonfly and damselfly diversity in West and Central Asia and show that future assessments of the threat status of freshwater taxa are of little use when climate change is not taken fully into account. It is recommended that the modelled future distributions are used to design a future-proof network of nature reserves encompassing the main freshwater systems. Furthermore, it is recommended that some basic monitoring system for freshwater diversity is established as currently, most data are still dependent on work by foreign visitors often collecting data as part of their holiday. Establishing a monitoring system will on a national scale allow governments to understand the impact of their policy and will on a regional scale allow them to detect if species are indeed able to colonize newly available habitats before their current habitats cease to exist.

ACK N O WLE D G E M ENTS
LM is supported by an F.R.S. -FNRS fellowship (Chargé de recherches). The compilation of the database of the dragonflies and damselflies of West Asia was made possible by NLBIF. We would also like to thank all individuals and organizations involved in the collection of the dragonfly distribution data used in the study. The authors declare that they have no conflicts of interest.

CO N FLI C T O F I NTE R E S T S TATE M E NT
The authors declare that they have no conflicts of interest.

PEER R E V I E W
The peer review history for this article is available at https:// www.webof scien ce.com/api/gatew ay/wos/peer-revie w/10.1111/ ddi.13704.

DATA AVA I L A B I L I T Y S TAT E M E N T
The