Two new species of the family Galumnellidae (Acari: Oribatida) from India

Two new galumnellid mites, Galumnella parageographica n. sp. and Porogalumnella microsetosa n. sp., are described from Indian soils. Galumnella parageographica n. sp. is morphologically most similar to G. geographica Mahunka, 1995 from Malaysia, from which it differs by tuberculated pteromorphs, the presence of a median pore on the notogaster and monodactylous legs. Porogalumnella microsetosa n. sp. is morphologically most similar to P. reducta Mahunka, 1995 from Malaysia, from which it differs by the presence of polygonal ornamentation in the anogenital region and nearly smooth sensilli. An identification key to the known species of Porogalumnella is provided.


INTRODUCTION
During taxonomic identification of oribatid mites (Acari: Oribatida) from India, we found two new species of the family Galumnellidae, one belonging to the genus Galumnella Berlese, 1916, the other to Porogalumnella Balogh, 1968. In this paper the new species are described and illustrated under the names Galumnella parageographica n. sp. and Porogalumnella microsetosa n. sp.
The genus Galumnella was proposed by Berlese (1916) with Galumnella paradoxa Berlese, 1916 as the type species. Currently, it comprises more than 20 species, which are collectively distributed in the tropic regions. The main characters of Galumnella are (summarized by Balogh and Balogh 2002; Ermilov and Anichkin 2011; including our additions and corrections): body surface with ornamentation (foveolate, reticulate, punctate, granulate); lamel-lar and sublamellar lines present (rarely sublamellar lines absent); sensilli long, setiform or with slightly dilated, pointed head, directed backwards; notogaster without porose areas (sometimes with pores); notogastral setae represented by 10 short setae or their alveoli; epimeral neotrichy absent; leg tarsi with one or three claws. Identification keys for the many known species of this genus were presented by Balogh and Balogh (2002) and Ermilov and Anichkin (2011).
The genus Porogalumnella was proposed by Balogh (1968) with Porogalumnella quadriporosa Balogh, 1968 as the type species. Currently, it comprises five species, which are collectively distributed in the tropic regions. The main characters of Porogalumnella are (summarized by Balakrishnan and Haq 1982;Balogh and Balogh 1992; including our additions and corrections): body surface with ornamentation (foveolate, reticulate, punctate, granulate); lamellar and sublamellar lines present; sensilli long or of medium size, with dilated, pointed head, directed backwards; notogaster with porose areas; notogastral setae represented by 10 short or long setae or their alveoli; epimeral neotrichy absent; leg tarsi with one or two claws. Below, we provide the first identification key to known species of this genus.

MATERIALS AND METHODS
Specimens of both new species were collected at the same locality: India, Arunachal Pradesh, Hunli vicinity, 28°19'32"N, 95°57'31"E, 1300 m a.s.l. They were extracted from soil by L. Dembický and O. Šauša on 01.VI.2012. Galumnella parageographica n. sp. is represented by the holotype (female) and one paratype (female); Porogalumnella microsetosa n. sp. is represented by the holotype (female) and four paratypes (three females, one male). The specimens were mounted in lactic acid on temporary cavity slides for measurement and illustration. All body measurements are presented in micrometers. The body length was measured in lateral view, from the tip of the rostrum to the posterior edge of the ventral plate. The notogastral width refers to the maximum width in dorsal aspect. Lengths of body setae were measured in lateral aspect. Formulae for leg setation are given in parentheses according to the sequence trochanter-femur-genu-tibia-tarsus (famulus included). Formulae for leg solenidia are given in square brackets according to the sequence genu-tibia-tarsus. General terminology used in this paper follows that of F. Grandjean (summarized by Norton and Behan-Pelletier 2009).
Type deposition -The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; the paratype is in the personal collection of the first author.
Etymology -The prefix para is Latin meaning "near" and refers the similarity between the new species and the species Galumnella geographica Mahunka, 1995. Remarks -Galumnella parageographica n. sp. is similar to G. geographica Mahunka, 1995 from Malaysia (see Mahunka 1995) in having setiform, barbed sensilli and polygonal ornamentation on the prodorsum, notogaster and anogenital region. However, it differs clearly from the latter by the having tubercles on the pteromorphs and a median notogastral pore (neither present in G. geographica) and monodactylous legs (tridactylous in G. geographica).
Integument - (Figure 2A, B, C, E). Body color dark-brown. Surface of prodorsum and notogaster with large round foveolae (diameter up to 8). Surface of anogenital region with polygonal ornamentation. Pteromorphs with polygonal foveolae. Genital plates with indistinct longitudinal lines.
Type deposition -The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; two paratype are deposited in the collection of the