Phenylalanine and tyrosine are incorporated specifically into annuloline in Lolium multiflorum. Tyramine, cinnamic acid, p-coumaric acid, and caffeic acid are shown to be intermediates in the biosynthesis. Methylation is shown to occur late in the biosynthetic sequence.
D. G. O'Donovan and H. Horan, J. Chem. Soc. C, 1971, 331 DOI: 10.1039/J39710000331
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