* Corresponding authors

^a CAS Key Laboratory of Tropical Marine Bio-resources and Ecology/Guangdong Key Laboratory of Marine Materia Medica/Innovation Academy of South China Sea Ecology and Environmental Engineering, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China
E-mail: wangjunfeng@scsio.ac.cn, yonghongliu@scsio.ac.cn

^b College of Food Science and Technology, Guangdong Provincial Key Laboratory of Aquatic Product Processing and Safety, Guangdong Province Engineering Laboratory for Marine Biological Products, Guangdong Provincial Engineering Technology Research Center of Seafood, Key Laboratory of Advanced Processing of Aquatic Product of Guangdong Higher Education Institution, Guangdong Ocean University, Zhanjiang 524088, China
E-mail: zyyang@gdou.edu.cn

^c University of Chinese Academy of Sciences, 19 Yuquan Road, Beijing 100049, China

^d Sanya Institute of Marine Ecology and Engineering, Yazhou Scientific Bay, Sanya 572000, China