Among the commonest and most abundant plant-remains in the Carmyllie and Cairnconon Beds of the Caledonian Lower Old Red Sandstone are branched linear axes about 2 mm. in width. These often occur in a fragmentary condition, but more connected portions demonstrating the morphology of the plant are also found. Though often associated with Parka and Nematophyton, the plant known as Zosterophyllum myretonianum is usually readily recognisable by its definite outline and uniform diameter, by its modes of branching, and by the remains of structure which it sometimes retains. As will be seen below, the axis of this plant was traversed by a single median strand that was composed of tracheides. Zosterophyllum is thus of special interest as the most ancient vascular plant known from British rocks. Some other plant-remains that may be confused with it have also been met with, though less commonly. These will be briefly considered, not merely for this reason, but on account of the very different and peculiar type of structure they exhibit.