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Re-Assessment of Late Campanian (Kirtlandian) Turtles from the Upper Cretaceous Fruitland and Kirtland Formations, San Juan Basin, New Mexico, USA

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Book cover Morphology and Evolution of Turtles

Abstract

The fossil turtles from the Upper Cretaceous Fruitland and Kirtland formations (late Campanian; Kirtlandian) have been known for more than 100 years. We re-assess and revise these Late Cretaceous testudine taxa from the San Juan Basin, New Mexico, USA, and discuss their biostratigraphic distribution. We recognize the following valid taxa as present: the bothremydid Chedighaii hutchisoni; the pleurosternid Compsemys sp.; the baenodds Denazinemys nodosa and Scabremys gen. nov., established for the distinct species S. ornata, previously included in Denazinemys; and Boremys grandis, though it is a rare taxon. The non-baenodd baenid Neurankylus baueri is recognized as a valid species. Two additional non-baenodd baenid taxa, Thescelus hemispherica and T. rapiens, are retained as distinct species and are not considered synonymous with T. insiliens. We also recognize a small indeterminate kinosternoid similar to that reported from the Campanian of Mexico. The two adocids Adocus bossi and A. kirtlandius are retained as distinct species. The nanhsiungchelyid Basilemys gaffneyi sp. nov. is established, whereas Basilemys nobilis is considered a nomen dubium because it lacks the diagnostic features that would allow referral to any known valid species. We recognize three trionychids: Aspideretoides austerus and A. robustus (new combination), and an unnamed plastominine. We synomize Aspideretoides fontanus and A. vorax with A. austerus. Aspideretoides ovatus is considered a subadult of A. robustus. The unnamed plastominine may represent a new genus and species. Turtles of the Fruitland-Kirtland formations resemble other late Campanian turtle assemblages from western North America, and are part of the characteristic vertebrate fauna of the Kirtlandian land-vertebrate age. The upper Fruitland and lower Kirtland formations (Hunter Wash local fauna) have greater turtle taxonomic diversity than the upper Kirtland Formation (Willow Wash local fauna). This apparent decrease in taxonomic diversity is interpreted as being real and reflects a shift in depositional (channel) environments to a more terrestrial one, a pattern which is seen in other North American Late Cretaceous settings.

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Acknowledgments

This study was made possible by the assistance and help of a number of individuals. For access to collections we thank Eugene S. Gaffney, Mark Norell, and Carl Mehling, (all American Museum of Natural History); John Bolt and William Simpson (both Field Museum of Natural History); Larry Martin and Desui Miao (both University of Kansas); Solwieg Stuenes (Paleontologiska Museet, University of Uppsala); and Michael Brett Surman (United States National Museum, Smithsonian). For discussions and verification of specimen identities we thank Don Brinkman (Royal Tyrrell Museum of Palaeontology); Walter Joyce (University of Tübingen), and Tyler Lyson (Yale University). For help in the field we thank John Burris (San Juan Community College), Mike Burns (University of Alberta), Denver Fowler (Montana State University), Kesler Randall (San Diego Natural History Museum), and Justin A. Spielmann (New Mexico Museum of Natural History and Science). Additional field assistance provided by independent individuals include Arjan Boere, James Hartley, James Nikas, Jim Murphy and Fred Widmann. Kevin Seymour (Royal Ontario Museum) kindly provided the information for ROM 864, and we thank him for these data. Donald Brinkman, Derek W. Larson (University of Toronto), and J. Howard Hutchison reviewed the manuscript, and we thank them for their valuable comments and suggestions. Finally, we thank Pat Hester and Sherrie Landon (Bureau of Land Management, Albuquerque and Farmington offices) for providing the necessary Paleontological Resource Permits for collecting specimens housed in the collections of the NMMNH and the SMP, and for their continuing support of our fieldwork.

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Correspondence to Robert M. Sullivan .

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Appendix

Appendix

List of all known previously referred, and newly referred, turtle specimens from the Fruitland and Kirtland formations, as of 2010. Order is alphabetic by (1) family/higher taxonomic category, (2) genus, (3) species, and (4) institution.

Current number

Previous number

Family

Genus

Species

Type status

Locality Number

Formation

Member

Description

USNM 8577

Adocidae

Adocus

bossi

paratype

Kirtland

Hunter Wash

partial carapace & plastron

USNM 8613

Adocidae

Adocus

bossi

holotype

Kirtland

Hunter Wash

partial carapace & nearly complete plastron

USNM 11082

Adocidae

Adocus

bossi

Kirtland

De-na-zin

carapace & plastron

USNM 11326

Adocidae

Adocus

bossi

Kirtland

Hunter Wash

incomplete carapace & plastron

USNM 12838

Adocidae

Adocus

bossi

Kirtland

Hunter Wash

carapace & plastron

USNM 12842

Adocidae

Adocus

bossi

Kirtland

Hunter Wash

carapace & plastron

USNM 12982

Adocidae

Adocus

bossi

Kirtland

Hunter Wash

carapace & plastron

USNM 12983

Adocidae

Adocus

bossi

Kirtland

Hunter Wash

carapace, plastron, & pelvis

USNM 8593

Adocidae

Adocus

kirtlandius

holotype

Kirtland

Hunter Wash

incomplete carapace & plastron

SMP VP-1725

Adocidae

Adocus

sp

313a

Kirtland

De-na-zin

carapace fragments

SMP VP-1280

Adocidae

Adocus

sp.

313a

Kirtland

De-na-zin

carapace & plastron fragments

SMP VP-1295

Adocidae

Adocus

sp.

350

Kirtland

De-na-zin

carapace & plastron fragments

SMP VP-1328

Adocidae

Adocus

sp.

386

Kirtland

Hunter Wash

peripheral carapace fragment

SMP VP-1330

Adocidae

Adocus

sp.

385

Kirtland

Hunter Wash

part of right carapace with peripherals & fragments

SMP VP-1331

Adocidae

Adocus

sp.

385

Kirtland

Hunter Wash

carapace/bridge fragment

SMP VP-1346

Adocidae

Adocus

sp.

373

Kirtland

Hunter Wash

carapace fragment with part of bridge

SMP VP-1350

Adocidae

Adocus

sp.

373

Kirtland

Hunter Wash

nuchal & anterior carapace fragments

SMP VP-1417

Adocidae

Adocus

sp.

388a

Kirtland

De-na-zin

carapace fragments

SMP VP-1482

Adocidae

Adocus

sp.

281

Kirtland

Hunter Wash

plastron & other fragments

SMP VP-1502

Adocidae

Adocus

sp.

283

Kirtland

Hunter Wash

carapace fragments

SMP VP-1504

Adocidae

Adocus

sp.

282

Kirtland

Hunter Wash

costals

SMP VP-1523

Adocidae

Adocus

sp.

281

Kirtland

Hunter Wash

peripheral of carapace

SMP VP-1544

Adocidae

Adocus

sp.

372

Kirtland

Hunter Wash

carapace fragment

SMP VP-1660

Adocidae

Adocus

sp.

283

Kirtland

Hunter Wash

carapace & plastron fragments

SMP VP-2032

Adocidae

Adocus

sp.

421

Kirtland

Hunter Wash

part of carapace

SMP VP-2033

Adocidae

Adocus

sp.

421

Kirtland

Hunter Wash

carapace fragments

SMP VP-2361

Adocidae

Adocus

sp.

451

Kirtland

Hunter Wash

carapace fragments & incomplete limb bone

SMP VP-2438

Adocidae

Adocus

sp.

435

Kirtland

Hunter Wash

carapace fragment

SMP VP-2439

Adocidae

Adocus

sp.

435

Kirtland

Hunter Wash

neural & other carapace fragments

SMP VP-2602

Adocidae

Adocus

sp.

319a

Kirtland

De-na-zin

two carapace fragments

SMP VP-2616

Adocidae

Adocus

sp.

319a

Kirtland

De-na-zin

carapace fragment

SMP VP-2818

Adocidae

Adocus

sp.

228

Kirtland

Hunter Wash

carapace neural

SMP VP-3264

Adocidae

Adocus

sp.

228

Kirtland

Hunter Wash

carapace fragments

SMP VP-3275

Adocidae

Adocus

sp.

463

Fruitland

Fossil Forest

carapace fragments

NMMNH P-22621

Adocidae

Adocus

sp.

1636

Kirtland

Hunter Wash

incomplete nuchal

NMMNH P-22700

Adocidae

Adocus

sp.

1623

Kirtland/Fruitland

unknown

6 carapace fragments

NMMNH P-22709

Adocidae

Adocus

sp.

1649

Kirtland

Hunter Wash

2 carapace fragments

NMMNH P-22747

Adocidae

Adocus

sp.

1674

Fruitland

Fossil Forest

carapace/plastron fragments

NMMNH P-26225

Adocidae

Adocus

sp.

3227

Kirtland

De-na-zin

incomplete neural

NMMNH P-26258

Adocidae

Adocus

sp.

3243

Kirtland

De-na-zin

carapace fragment

NMMNH P-26262

Adocidae

Adocus

sp.

3245

Kirtland

De-na-zin

incomplete costal

NMMNH P-27427

Adocidae

Adocus

sp.

1708

Kirtland

Hunter Wash

numerous shell fragments

NMMNH P-27463

Adocidae

Adocus

sp.

3875

Kirtland

Hunter Wash

23 carapace/plastron fragments

NMMNH P-27601

Adocidae

Adocus

sp.

4015

Fruitland

Fossil Forest

carapace/plastron fragments

NMMNH P-27629

Adocidae

Adocus

sp.

4015

Fruitland

Fossil Forest

carapace fragments

NMMNH P-27760

Adocidae

Adocus

sp.

4015

Fruitland

Fossil Forest

carapace/plastron fragments

NMMNH P-27774

Adocidae

Adocus

sp.

4015

Fruitland

Fossil Forest

carapace fragments & neural

NMMNH P-27926

Adocidae

Adocus

sp.

1688

Kirtland

Hunter Wash

shell fragments

NMMNH P-30759

Adocidae

Adocus

sp.

3117

Kirtland

Hunter Wash

miscellaneous shell fragments

NMMNH P-32871

Adocidae

Adocus

sp.

2387

Fruitland

Fossil Forest

6 shell fragments

NMMNH P-32964

Adocidae

Adocus

sp.

2387

Fruitland

Fossil Forest

13 fragments

NMMNH P-38403

Adocidae

Adocus

sp.

4010

Fruitland

Fossil Forest

entoplaston

NMMNH P-41022

Adocidae

Adocus

sp.

5400

Kirtland

De-na-zin

carapace fragments

NMMNH P-41023

Adocidae

Adocus

sp.

5400

Kirtland

De-na-zin

15 carapace/plastron fragments

NMMNH P-49867

UA 5572

Adocidae

Adocus

sp.

6203

Fruitland?

Fossil Forest?

nearly complete plastron

NMMNH P-22537

Adocidae

Adocus

sp.

1582

Kirtland

Hunter Wash

shell fragment

NMMNH P-22543

Adocidae

Adocus

sp.

1584

Kirtland

Hunter Wash

shell fragment

NMMNH P-22652

Adocidae

Adocus

sp.

1672

Fruitland

Fossil Forest

3 shell fragments

NMMNH P-22653

Adocidae

Adocus

sp.

1672

Fruitland

Fossil Forest

shell fragment

NMMNH P-22733

Adocidae

Adocus

sp.

1668

Kirtland

Hunter Wash

shell fragments

NMMNH P-22740

Adocidae

Adocus

sp.

1674

Fruitland

Fossil Forest

carapace fragments

NMMNH P-22750

Adocidae

Adocus

sp.

1674

Fruitland

Fossil Forest

incomplete costals

NMMNH P-22822

Adocidae

Adocus

sp.

1706

Fruitland

Fossil Forest

2 shell fragments

NMMNH P-27883

Adocidae

Adocus

sp.

3227

Kirtland

De-na-zin

costal, plastron, peripheral, & other fragments

NMMNH P-28327

Adocidae

Adocus

sp.

1882

Fruitland

Fossil Forest

2 carapace fragments

NMMNH P-30069

Adocidae

Adocus

sp.

4267

Fruitland

Fossil Forest

numerous shell fragments

NMMNH P-30361

Adocidae

Adocus

sp.

3490

Kirtland

Hunter Wash

numerous carapace/plastron fragments

NMMNH P-32850

Adocidae

Adocus

sp.

4544

Fruitland

Fossil Forest

14 costal fragments

NMMNH P-49724

UA 13483

Adocidae

Adocus

sp.

6333

Kirtland?

Hunter Wash?

shell fragment

NMMNH P-49725

UA 14413

Adocidae

Adocus

sp.

6333

Kirtland?

Hunter Wash?

4 shell fragments

NMMNH P-49826

UA 14404

Adocidae

Adocus

sp.

6559

Kirtland?

Hunter Wash?

2 shell fragments

NMMNH P-49831

UA 14411

Adocidae

Adocus

sp.

6622

Kirtland?

De-na-zin?

shell fragment

NMMNH P-49849

UA 14402

Adocidae

Adocus

sp.

6547

Kirtland?

Hunter Wash?

2 shell fragments

NMMNH P-49851

UA 14405

Adocidae

Adocus

sp.

6551

Kirtland?

De-na-zin?

carapace fragment

NMMNH P-49854

UA 14401

Adocidae

Adocus

sp.

6552

Kirtland?

De-na-zin?

shell fragment

NMMNH P-49859

UA 14403

Adocidae

Adocus

sp.

6558

Kirtland?

Hunter Wash?

2 shell fragments

NMMNH P-49866

UA 5575

Adocidae

Adocus

sp.

6203

Fruitland?

Fossil Forest?

plastron & carapace fragments

NMMNH P-49895

UA 14406

Adocidae

Adocus

sp.

6542

Kirtland?

Hunter Wash?

shell fragment

NMMNH P-49926

UA 8559

Adocidae

Adocus

sp.

6272

Kirtland?

Hunter Wash?

shell fragment

NMMNH P-49927

UA 8560

Adocidae

Adocus

sp.

6272

Kirtland?

Hunter Wash?

shell fragment

NMMNH P-50625

UA 14407

Adocidae

Adocus

sp.

6558

Kirtland?

Hunter Wash?

shell fragment

NMMNH P-49846

UA 14426

Adocidae

Adocus

sp.

6544

Kirtland?

Hunter Wash?

2 shell fragment

USNM 8654

Adocidae

Adocus

sp.

Kirtland

Hunter Wash

partial plastron

SMP VP-1098

Adocidae

cf. Adocus

sp.

361

Kirtland

De-na-zin

carapace & plastron fragments

SMP VP-1294

Adocidae

cf. Adocus

sp.

372

Kirtland

Hunter Wash

carapace & plastron fragments from multiple individuals

SMP VP-1477

Adocidae

cf. Adocus

sp.

365

Kirtland

Hunter Wash

carapace fragment

SMP VP-1666

Adocidae

cf. Adocus

sp.

398

Kirtland

Hunter Wash

carapace & plastron fragments

SMP VP-1843

Adocidae

cf. Adocus

sp.

319a

Kirtland

De-na-zin

plastron fragment

SMP VP-2096

Adocidae

cf. Adocus

sp.

389a

Kirtland

De-na-zin

incomplete plastron

SMP VP-2382

Adocidae

cf. Adocus

sp.

382

Kirtland

De-na-zin

carapace fragment

SMP VP-2578

Adocidae

cf. Adocus

sp.

450

Fruitland

Fossil Forest

neural

SMP VP-2841

Adocidae

cf. Adocus

sp.

450

Fruitland

Fossil Forest

carapace fragment & associated indeterminate bone material

SMP VP-2856

Adocidae

cf. Adocus

sp.

450

Fruitland

Fossil Forest

incomplete costal & other carapace fragments

NMMNH P-28322

Adocidae

cf. Adocus

sp.

1882

Fruitland

Fossil Forest

numerous carapace fragments

NMMNH P-44902

Adocidae

cf. Adocus

sp.

3517

Kirtland

Hunter Wash

carapace fragments

NMMNH P-22756

Adocidae

?Adocus

sp.

1678

Fruitland

Fossil Forest

various shell fragments

NMMNH P-27781

Adocidae

?Adocus

sp.

4015

Fruitland

Fossil Forest

numerous carapace/plastron fragments

SMP VP-1444

Adocidae

?Adocus

sp.

281

Kirtland

Hunter Wash

fragments of carapace

SMP VP-1956

Adocidae

?Adocus

sp.

382

Kirtland

De-na-zin

neural

SMP VP-2018

Adocidae

?Adocus

sp.

420a

Kirtland

De-na-zin

plastron fragment

SMP VP-2491

Adocidae

?Adocus

sp.

389a

Kirtland

De-na-zin

carapace fragment

SMP VP-2802

Adocidae

?Adocus

sp.

450

Fruitland

Fossil Forest

costal, marginal, & other carapace fragments

USNM 12979

Baenidae

Boremys

grandis

holotype

Kirtland

De-na-zin

carapace & plastron

USNM 12978

Baenidae

Boremys

sp.

Kirtland

Hunter Wash?

partial carapace & plastron

SMP VP-1565

Baenidae

cf. Boremys

sp.

391

Kirtland

Hunter Wash

plastron fragment

FMNH PR-2250

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

nearly complete carapace & plastron

NMMNH P-22564

Baenidae

Denazinemys

nodosa

1602

Kirtland

Farmington

2 carapace fragments

NMMNH P-22716

Baenidae

Denazinemys

nodosa

1654

Kirtland

Hunter Wash

2 shell fragments

NMMNH P-22790

Baenidae

Denazinemys

nodosa

1696

Fruitland

Fossil Forest

numerous shell fragments

NMMNH P-22881

Baenidae

Denazinemys

nodosa

1755

Kirtland

Hunter Wash

numerous carapace & bone fragments

NMMNH P-25075

Baenidae

Denazinemys

nodosa

1674

Fruitland

Fossil Forest

nearly complete

NMMNH P-25087

Baenidae

Denazinemys

nodosa

3117

Kirtland

Hunter Wash

complete carapace & plastron

NMMNH P-27732

Baenidae

Denazinemys

nodosa

4015

Fruitland

Fossil Forest

carapace fragments

NMMNH P-27830

Baenidae

Denazinemys

nodosa

4015

Fruitland

Fossil Forest

partial carapace & plastron

NMMNH P-32890

Baenidae

Denazinemys

nodosa

4010

Kirtland

Hunter Wash

incomplete carapace & fragments

NMMNH P-37763

Baenidae

Denazinemys

nodosa

5163

Kirtland

Hunter Wash

numerous carapace & plastron fragments

NMMNH P-39633

Baenidae

Denazinemys

nodosa

4275

Kirtland

Hunter Wash

numerous carapace & plastron fragments

NMMNH P-49858

UA 14394

Baenidae

Denazinemys

nodosa

6558

Kirtland?

Hunter Wash?

shell fragment

NMMNH P-49896

UA 8826

Baenidae

Denazinemys

nodosa

6539

Kirtland?

Hunter Wash?

carapace fragment

NMMNH P-26254

Baenidae

Denazinemys

nodosa

3226

Kirtland

De-na-zin

carapace fragment

NMMNH P-26255

Baenidae

Denazinemys

nodosa

3226

Kirtland

De-na-zin

carapace fragment

NMMNH P-26298

Baenidae

Denazinemys

nodosa

3169

Kirtland

De-na-zin

numerous carapace fragments

NMMNH P-27278

Baenidae

Denazinemys

nodosa

3530

Kirtland

De-na-zin

2 carapace fragments

NMMNH P-27439

Baenidae

Denazinemys

nodosa

3531

Kirtland

De-na-zin

neural part of carapace

NMMNH P-33026

Baenidae

Denazinemys

nodosa

4275

Kirtland

Hunter Wash

12 carapace fragments

NMMNH P-33028

Baenidae

Denazinemys

nodosa

4275

Kirtland

Hunter Wash

neural fragment

NMMNH P-41229

Baenidae

Denazinemys

nodosa

3506

Kirtland

Hunter Wash

nearly complete carapace & plastron

NMMNH P-22636

Baenidae

Denazinemys

nodosa

1640

Kirtland

Hunter Wash

carapace fragments

NMMNH P-22643

Baenidae

Denazinemys

nodosa

1606

Fruitland

Fossil Forest

carapace fragments

NMMNH P-28323

Baenidae

Denazinemys

nodosa

1882

Fruitland

Fossil Forest

numerous carapace fragments

NMMNH P-32867

Baenidae

Denazinemys

nodosa

2387

Fruitland

Fossil Forest

5 carapace fragments

PMU.R 11

Exemplar 1

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

carapace, plastron, pelvis, & other postcrania

PMU.R 12

Exemplar 2

Baenidae

Denazinemys

nodosa

Kirtland

De-na-zin

nearly complete carapace & plastron

PMU.R 13

Exemplar 3

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

incomplete carapace & [plastron

PMU.R 14

Exemplar 4

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

incomplete carapace & plastron

PMU.R unnumbered

Exemplar 5

Baenidae

Denazinemys

nodosa

unknown

unknown

incomplete carapace & plastron

 PMU.R unnumbered

Exemplar 6

Baenidae

Denazinemys

nodosa

Kirtland

De-na-zin

incomplete carapace & plastron

PMU.R 15

Exemplar 7

Baenidae

Denazinemys

nodosa

unknown

unknown

incomplete carapace

 PMU.R unnumbered

Exemplar 8

Baenidae

Denazinemys

nodosa

Kirtland

De-na-zin

incomplete plastron & other shell material

PMU.R 16

Exemplar 9

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

incomplete carapace

PMU.R 17

Exemplar 10

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

incomplete carapace & nearly complete plastron

PMU.R 18

Exemplar 11

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

nearly complete carapace & plastron

PMU.R 19

Exemplar 12

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

incomplete plastron

PMU.R 20

Exemplar 13

Baenidae

Denazinemys

nodosa

unknown

unknown

incomplete carapace & plastron

 PMU.R unnumbered

Exemplar 14

Baenidae

Denazinemys

nodosa

unknown

unknown

shell fragments

 PMU.R unnumbered

Exemplar 15

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

carapace & plastron fragments

 PMU.R unnumbered

Exemplar 16

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

carapace fragments

 PMU.R unnumbered

Exemplar 17

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

shell fragments

SMP VP-1284

Baenidae

Denazinemys

nodosa

350

Kirtland

De-na-zin

?carapace fragment

SMP VP-1287

Baenidae

Denazinemys

nodosa

350

Kirtland

De-na-zin

numerous plastron & carapace fragments

SMP VP-1290

Baenidae

Denazinemys

nodosa

360a

Kirtland

De-na-zin

carapace fragments

SMP VP-1323

Baenidae

Denazinemys

nodosa

319a

Kirtland

De-na-zin

carapace fragment

SMP VP-1403

Baenidae

Denazinemys

nodosa

388a

Kirtland

De-na-zin

carapace & associated fragments

SMP VP-1869

Baenidae

Denazinemys

nodosa

382

Kirtland

De-na-zin

complete carapace & plastron

SMP VP-2072

Baenidae

Denazinemys

nodosa

376a

Kirtland

De-na-zin

nearly complete carapace & plaston

SMP VP-2495

Baenidae

Denazinemys

nodosa

389a

Kirtland

De-na-zin

carapace fragment

SMP VP-830

Baenidae

Denazinemys

nodosa

316

Kirtland

Hunter Wash

incomplete plastron with left bridge & numerous carapace fragments

SMP VP-1348

Baenidae

Denazinemys

nodosa

373

Kirtland

Hunter Wash

carapace fragments

SMP VP-1469

Baenidae

Denazinemys

nodosa

228

Kirtland

Hunter Wash

anterior part of plastron (intergular, gular, & humeral)

SMP VP-1480

Baenidae

Denazinemys

nodosa

281

Kirtland

Hunter Wash

carapace fragment

SMP VP-1506

Baenidae

Denazinemys

nodosa

228

Kirtland

Hunter Wash

right plastron fragment (intergular, gular, & humeral)

SMP VP-1524

Baenidae

Denazinemys

nodosa

281

Kirtland

Hunter Wash

carapace fragment

SMP VP-1665

Baenidae

Denazinemys

nodosa

398

Kirtland

Hunter Wash

medial part of carapace

SMP VP-1668

Baenidae

Denazinemys

nodosa

398

Kirtland

Hunter Wash

complete plastron, partial carapace, & fragments

SMP VP-1948

Baenidae

Denazinemys

nodosa

228

Kirtland

Hunter Wash

four carapace fragments

SMP VP-1986

Baenidae

Denazinemys

nodosa

419

Kirtland

Hunter Wash

incomplete plastron & carapace

SMP VP-2529

Baenidae

Denazinemys

nodosa

452

Kirtland

Hunter Wash

carapace fragment

SMP VP-2828

Baenidae

Denazinemys

nodosa

349

Kirtland

Hunter Wash

incomplete plastron

SMP VP-3255

Baenidae

Denazinemys

nodosa

398

Kirtland

Hunter Wash

plastron fragment, including part of xiphiplastron & bridge

USNM 8345

Baenidae

Denazinemys

nodosa

holotype

Kirtland

De-na-zin

carapace & plastron

USNM 8603

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

median part of carapace

USNM 8622

Baenidae

Denazinemys

nodosa

Fruitland

Fossil Forest

median part of carapace

USNM 11323

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

carapace & plastron

USNM 11327

Baenidae

Denazinemys

nodosa

Fruitland

Fossil Forest

crushed shell

USNM 12819

Baenidae

Denazinemys

nodosa

Kirtland

Hunter Wash

carapace & incomplete plastron

USNM 12834

Baenidae

Denazinemys

nodosa

Kirtland

De-na-zin

incomplete carapace & plastron

NMMNH P-49945

UA 5573

Baenidae

cf. Denazinemys

nodosa

6205

Kirtland?

Hunter Wash?

10 plastron & associated fragments

SMP VP-1704

Baenidae

cf. Denazinemys

nodosa

409

Kirtland

Hunter Wash

partial carapace & plastron

SMP VP-2853

Baenidae

cf. Denazinemys

nodosa

450

Fruitland

Fossil Forest

carapace fragments

SMP VP-3273

Baenidae

cf. Denazinemys

nodosa

372

Kirtland

Hunter Wash

carapace fragments

SMP VP-1610

Baenidae

?Denazinemys

nodosa

398

Kirtland

Hunter Wash

numerous carapace & plastron fragments

SMP VP-2751

Baenidae

?Denazinemys

nodosa

228

Kirtland

Hunter Wash

carapace fragments

NMMNH P-22698

Baenidae

Neurankylus

baueri

1616

Kirtland

Hunter Wash

carapace fragments

NMMNH P-27988

 

Baenidae

Neurankylus

baueri

3117

Kirtland

Hunter Wash

carapace & plastron sections

NMMNH P-22745

Baenidae

Neurankylus

baueri

1674

Fruitland

Fossil Forest

various shell fragments

NMMNH P-27788

Baenidae

Neurankylus

baueri

4015

Fruitland

Fossil Forest

numerous shell fragments

PMU.R 24

Exemplar 1

Baenidae

Neurankylus

baueri

Kirtland

Hunter Wash

nearly complete carapace, plastron, axial, & appendicular skeleton

PMU.R 25

Exemplar 2

Baenidae

Neurankylus

baueri

Kirtland

Hunter Wash

nearly complete carapace & incomplete plastron

PMU.R 26

Exemplar 3

Baenidae

Neurankylus

baueri

Kirtland

Hunter Wash?

incomplete carapace, plastron, & pelvic girdle

PMU.R 27

Exemplar 4

Baenidae

Neurankylus

baueri

Kirtland

Hunter Wash

incomplete carapace

SMP VP-1503

Baenidae

Neurankylus

baueri

283

Kirtland

Hunter Wash

plastron fragments

SMP VP-2379

Baenidae

Neurankylus

baueri

389a

Kirtland

De-na-zin

carapace with associated fragments & plastron

USNM 8344

Baenidae

Neurankylus

baueri

holotype

Kirtland

Hunter Wash

nearly complete carapace & plastron

USNM 8531

Baenidae

Neurankylus

baueri

hypotype

Kirtland

Hunter Wash

carapace & plastron

NMMNH P-22749

Baenidae

Neurankylus

cf. N. baueri

1674

Fruitland

Fossil Forest

shell fragment

NMMNH P-27776

Baenidae

Neurankylus

cf. N. baueri

4015

Fruitland

Fossil Forest

numerous shell fragments

NMMNH P-22655

Baenidae

Neurankylus

cf. N. baueri

1672

Fruitland

Fossil Forest

3 shell fragments

NMMNH P-22713

Baenidae

Neurankylus

cf. N. baueri

1652

Kirtland

Hunter Wash

various shell & bone fragments

NMMNH P-22792

Baenidae

Neurankylus

cf. N. baueri

1697

Kirtland

Hunter Wash

incomplete plastron & various fragments

NMMNH P-28850

Baenidae

?Neurankylus

cf. N. baueri

3117

Kirtland

Hunter Wash

carapace sections

PMU.R 28

Baenidae

Neurankylus

cf. N. baueri

Kirtland

De-na-zin?

incomplete carapace & plastron

ROM 864

Baenidae

Neurankylus

cf. N. baueri

 

Kirtland

Hunter Wash

incomplete carapace & plastron

SMP VP-1854

Baenidae

Neurankylus

cf. N. baueri

283

Kirtland

Hunter Wash

portion of plastron

SMP VP-1931

Baenidae

Neurankylus

cf. N. baueri

228

Kirtland

Hunter Wash

partial carapace

SMP VP-2102

Baenidae

Neurankylus

cf. N. baueri

421

Kirtland

Hunter Wash

incomplete carapace & nearly complete plastron

USNM 8621

Baenidae

Neurankylus

cf. N. baueri

Kirtland

Hunter Wash

posterior carapace

USNM 13228

Baenidae

Neurankylus

cf. N. baueri

Kirtland

Hunter Wash

carapace & plastron

USNM 11083

Baenidae

Scabremys

ornata

Kirtland

De-na-zin

carapace & plastron

USNM 12821

Baenidae

Scabremys

ornata

Kirtland

Hunter Wash

partial carapace & plastron

USNM 13229

Baenidae

Scabremys

ornata

holotype

Kirtland

Hunter Wash

carapace & plastron

AMNH 6066

Baenidae

Thescelus

rapiens

holotype

Kirtland

De-na-zin

incomplete carapace & plastron

SMP VP-2100

Baenidae

Thescelus

rapiens

421

Kirtland

Hunter Wash

plastron & carapace

PMU.R 22

Baenidae

Thescelus

cf. T. rapiens

Kirtland

Hunter Wash

incomplete carapace & complete plastron

SMP VP-1553

Baenidae

Thescelus

cf. T. rapiens

393

Kirtland

De-na-zin

carapace fragments & associated fragments

USNM 12818

Baenidae

Thescelus

hemispherica

holotype

Kirtland

Hunter Wash

carapace & plastron

PMU.R 23

Baenidae

Thescelus

cf. T. hemispherica

Kirtland

Hunter Wash

incomplete carapace & plastron

SMP VP-2168

Baenidae

Thescelus

sp.

421

Kirtland

Hunter Wash

nearly complete carapace & plastron

NMMNH P-49940

UA 5586

Baenidae

Baenidae indet.

Baenidae indet.

6205

Kirtland?

Hunter Wash?

plastron fragment

SMP VP-1845

Baenidae

Baenidae indet.

Baenidae indet.

319a

Kirtland

De-na-zin

carapace section

SMP VP-2055

Baenidae

Baenidae indet.

Baenidae indet.

421

Kirtland

Hunter Wash

costal fragment

SMP VP-2600

Baenidae

Baenidae indet.

Baenidae indet.

319a

Kirtland

De-na-zin

incomplete plastral bridge

SMP VP-3257

Baenidae

Baenidae indet.

Baenidae indet.

313a

Kirtland

De-na-zin

plastron fragment

SMP VP-3258

Baenidae

Baenidae indet.

Baenidae indet.

350

Kirtland

De-na-zin

carapace fragment

NMMNH P-22793

Baenidae

Baenidae indet.

Baenidae indet.

1698

Fruitland

Fossil Forest

various shell and bone fragments

NMMNH P-22796

Baenidae

Baenidae indet.

Baenidae indet.

1699

Kirtland

Hunter Wash

shell fragment

NMMNH P-22841

Baenidae

Baenidae indet.

Baenidae indet.

1721

Kirtland

De-na-zin

carapace/plastron fragments

NMMNH P-27632

Baenidae

Baenidae indet.

Baenidae indet.

4015

Fruitland

Fossil Forest

numerous shell fragments

NMMNH P-27759

Baenidae

Baenidae indet.

Baenidae indet.

4015

Fruitland

Fossil Forest

4 carapace fragments

NMMNH P-27762

Baenidae

Baenidae indet.

Baenidae indet.

4015

Fruitland

Fossil Forest

plastron/carapace fragments

NMMNH P-33024

Baenidae

Baenidae indet.

Baenidae indet.

4507

Kirtland

De-na-zin

incomplete innominate

NMMNH P-38401

Baenidae

Baenidae indet.

Baenidae indet.

4010

Kirtland

Hunter Wash

incomplete carapace

SMP VP-1154

Baenidae

Baenidae indet.

Baenidae indet.

372

Kirtland

Hunter Wash

costal plate

SMP VP-1387

Baenidae

Baenidae indet.

Baenidae indet.

391

Kirtland

Hunter Wash

carapace fragments

SMP VP-1399

Baenidae

Baenidae indet.

Baenidae indet.

388a

Kirtland

De-na-zin

right xiphiplastron

SMP VP-1470

Baenidae

Baenidae indet.

Baenidae indet.

281

Kirtland

Hunter Wash

carapace fragment

SMP VP-1507

Baenidae

Baenidae indet.

Baenidae indet.

228

Kirtland

Hunter Wash

part of carapace

SMP VP-1661

Baenidae

Baenidae indet.

Baenidae indet.

398

Kirtland

Hunter Wash

carapace & plastron fragments

SMP VP-1669

Baenidae

Baenidae indet.

Baenidae indet.

398

Kirtland

Hunter Wash

incomplete carapace & associated fragments

SMP VP-1791

Baenidae

Baenidae indet.

Baenidae indet.

281

Kirtland

Hunter Wash

xiphiplastron & other plastron fragments

SMP VP-1798

Baenidae

Baenidae indet.

Baenidae indet.

228

Kirtland

Hunter Wash

dorsal vertebra

SMP VP-1946

Baenidae

Baenidae indet.

Baenidae indet.

228

Kirtland

Hunter Wash

costal

SMP VP-2014

Baenidae

Baenidae indet.

Baenidae indet.

388a

Kirtland

De-na-zin

complete lower jaw

SMP VP-2644

Baenidae

Baenidae indet.

Baenidae indet.

450

Fruitland

Fossil Forest

nearly complete femur

SMP VP-2706

Baenidae

Baenidae indet.

Baenidae indet.

450

Fruitland

Fossil Forest

incomplete costal & plastron fragments

SMP VP-2721

Baenidae

Baenidae indet.

Baenidae indet.

450

Fruitland

Fossil Forest

carapace & plastron fragments, plus associated indet. bone material

USNM 8614

Baenidae

Baenidae indet.

Baenidae indet.

Kirtland

Hunter Wash

anterior half of plastron

USNM 8623

Baenidae

Baenidae indet.

Baenidae indet.

Kirtland

Hunter Wash

anterior half of plastron

USNM 8624

Baenidae

Baenidae indet.

Baenidae indet.

Kirtland

Hunter Wash

partial plastron

NMMNH P-22878

?Baenidae

?Baenidae indet.

?Baenidae indet.

1754

Kirtland

Hunter Wash

bone scrap

NMMNH P-32854

?Baenidae

?Baenidae indet.

?Baenidae indet.

4541

Kirtland

De-na-zin

15 carapace/plastron fragments

SMP VP-1491

?Baenidae

?Baenidae indet.

?Baenidae indet.

309a

Kirtland

De-na-zin

posterior marginal?

SMP VP-1670

?Baenidae

?Baenidae indet.

?Baenidae indet.

399

Kirtland

Hunter Wash

nuchal & plastron fragments

SMP VP-2358

?Baenidae

?Baenidae indet.

?Baenidae indet.

281

Kirtland

Hunter Wash

plastron fragments

SMP VP-2605

?Baenidae

?Baenidae indet.

?Baenidae indet.

319a

Kirtland

De-na-zin

neural

SMP VP-2820

?Baenidae

?Baenidae indet.

?Baenidae indet.

228

Kirtland

Hunter Wash

carapace neural

NMMNH P-22813

?Baenidae

?Baenidae indet.

?Baenidae indet.

1702

Fruitland

Fossil Forest

?plastron section & various carapace/plastron fragments

SMP VP-3389

?Baenidae indet.

?Baenidae indet.

?Baenidae indet.

461

Kirtland

Hunter Wash

marginal/carapace fragment

SMP VP-1907

Kinosternoidea indet.

Kinosternoidea indet.

Kinosternoidea indet.

350

Kirtland

De-na-zin

part of left hypoplastron

SMP VP-2004

Kinosternoidea indet.

Kinosternoidea indet.

Kinosternoidea indet.

350

Kirtland

De-na-zin

two costal fragments & right hypoplastron

SMP VP-2009

Kinosternoidea indet.

Kinosternoidea indet.

Kinosternoidea indet.

350

Kirtland

De-na-zin

right xiphiplastron

SMP VP-2533

Kinosternoidea indet.

Kinosternoidea indet.

Kinosternoidea indet.

350

Kirtland

De-na-zin

right costal (8th?)

SMP VP-3274

Kinosternoidea indet.

Kinosternoidea indet.

Kinosternoidea indet.

350

Kirtland

De-na-zin

left costal (2nd?)

NMMNH P-49785

UA 14836

Kinosternoidea indet.

Kinosternoidea indet.

Kinosternoidea indet.

6340

Kirtland?

De-na-zin?

peripheral

USNM 11084

Nanhsiungchelyidae

Basilemys

gaffneyi

holotype

Kirtland

Hunter Wash

carapace & plastron

NMMNH P-27613

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

4015

Fruitland

Fossil Forest

carapace fragments

NMMNH P-27623

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

4015

Fruitland

Fossil Forest

peripheral fragment

NMMNH P-27630

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

4015

Fruitland

Fossil Forest

numerous fragments

NMMNH P-27775

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

4015

Fruitland

Fossil Forest

plastron section & 3 fragments

NMMNH P-28935

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

3510

Kirtland

Hunter Wash

carapace fragments

NMMNH P-28952

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

3496

Kirtland

Hunter Wash

peripheral fragments

NMMNH P-22651

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

1672

Fruitland

Fossil Forest

shell fragment

NMMNH P-22742

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

1674

Fruitland

Fossil Forest

shell fragments

NMMNH P-22744

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

1674

Fruitland

Fossil Forest

various shell fragments

NMMNH P-22871

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

1746

Kirtland

Hunter Wash

various shell & bone fragments

NMMNH P-22935

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

1791

Kirtland

Hunter Wash

carapace fragment

NMMNH P-22947

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

1777

Kirtland

Hunter Wash

limb osteoderm

NMMNH P-27942

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

1586

Kirtland

Hunter Wash

carapace fragments

NMMNH P-32828

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

4010

Kirtland

Hunter Wash

limb osteoderm

NMMNH P-44517

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

4000

Kirtland?

Hunter Wash?

carapace fragments

PMU.R 29

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

Kirtland

Hunter Wash

nearly complete carapace & plastron

SMP VP-1106

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

316

Kirtland

Hunter Wash

incomplete humerus & femur, plus other indet. fragments

SMP VP-1382

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

361

Kirtland

De-na-zin

carapace fragment

SMP VP-1475

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

365

Kirtland

Hunter Wash

edge of carapace

SMP VP-1510

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

228

Kirtland

Hunter Wash

plastron fragment

SMP VP-1522

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

281

Kirtland

Hunter Wash

carapace fragments

SMP VP-1527

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

281

Kirtland

Hunter Wash

carapace fragments

SMP VP-1566

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

281

Kirtland

Hunter Wash

carapace fragments

SMP VP-1652

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

281

Kirtland

Hunter Wash

carapace fragment

SMP VP-1653

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

281

Kirtland

Hunter Wash

carapace/plastron fragments

SMP VP-1678

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

400

Kirtland

Hunter Wash

partial right humerus

SMP VP-1852

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

228

Kirtland

Hunter Wash

limb osteoderm

SMP VP-2153

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

281

Kirtland

Hunter Wash

carapace fragment

SMP VP-2359

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

281

Kirtland

Hunter Wash

carapace fragment

SMP VP-2797

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

450

Fruitland

Fossil Forest

epiplastron

SMP VP-3366

Nanhsiungchelyidae

Basilemys

cf. B. gaffneyi

450

Fruitland

Fossil Forest

partial plastron, fragments of femur, pelvic girdle, & carapace/plastron

SMP VP-1146

Nanhsiungchelyidae

cf. Basilemys

cf. B. gaffneyi

365

Kirtland

Hunter Wash

terminal phalanx

SMP VP-1158

Nanhsiungchelyidae

cf. Basilemys

cf. B. gaffneyi

228

Kirtland

Hunter Wash

incomplete humerus

NMMNH P-22568

Pleurosternidae

Compsemys

sp.

1464

Kirtland

Hunter Wash

2 carapace/plastron fragments

NMMNH P-22741

Pleurosternidae

Compsemys

sp.

1674

Fruitland

Fossil Forest

plastron fragment

NMMNH P-22809

Pleurosternidae

Compsemys

sp.

1700

Fruitland

Fossil Forest

3 shell fragments

NMMNH P-28326

Pleurosternidae

Compsemys

sp.

1882

Fruitland

Fossil Forest

2 shell fragments

NMMNH P-33021

Pleurosternidae

Compsemys

sp.

4570

Kirtland

De-na-zin

4 carapace fragments

NMMNH P-40512

Pleurosternidae

Compsemys

sp.

3488

Kirtland

De-na-zin

incomplete carapace & plastron

NMMNH P-49819

UA 14391

Pleurosternidae

Compsemys

sp.

6551

Kirtland?

De-na-zin?

carapace fragment

NMMNH P-49821

UA 14388

Pleurosternidae

Compsemys

sp.

6554

Kirtland?

De-na-zin?

entoplaston

NMMNH P-49827

UA 14393

Pleurosternidae

Compsemys

sp.

6559

Kirtland?

Hunter Wash?

4 shell fragment

NMMNH P-49844

UA 14392

Pleurosternidae

Compsemys

sp.

6542

Kirtland?

Hunter Wash?

plastron fragment

SMP VP-1892

Pleurosternidae

Compsemys

sp.

350

Kirtland

De-na-zin

peripheral

SMP VP-1904

Pleurosternidae

Compsemys

sp.

350

Kirtland

De-na-zin

carapace fragment

SMP VP-1915

Pleurosternidae

Compsemys

sp.

350

Kirtland

De-na-zin

five carapace fragments

SMP VP-1916

Pleurosternidae

Compsemys

sp.

350

Kirtland

De-na-zin

carapace fragment

SMP VP-2002

Pleurosternidae

Compsemys

sp.

350

Kirtland

De-na-zin

costal fragment

SMP VP-2003

Pleurosternidae

Compsemys

sp.

350

Kirtland

De-na-zin

five costal fragments

SMP VP-2006

Pleurosternidae

Compsemys

sp.

350

Kirtland

De-na-zin

four plastron fragments

SMP VP-2201

Pleurosternidae

Compsemys

sp.

319a

Kirtland

De-na-zin

peripheral

SMP VP-2648

Pleurosternidae

Compsemys

sp.

350

Kirtland

De-na-zin

carapace fragments

SMP VP-2630

Pleurosternidae

?Compsemys

sp.

313a

Kirtland

De-na-zin

incomplete costal

NMMNH P-26230

Trionychidae

Aspideretoides

austerus

3227

Kirtland

De-na-zin

costal fragments & neural

NMMNH P-27847

Trionychidae

Aspideretoides

austerus

3226

Kirtland

De-na-zin

incomplete costals, xiphiplastron, hyoplastron, & girdle

SMP VP-811

Trionychidae

Aspideretoides

austerus

210

Kirtland

De-na-zin

several pieces of plastron and carapace

SMP VP-1364

Trionychidae

Aspideretoides

austerus

389a

Kirtland

De-na-zin

hypoplastron & other plastral elements

SMP VP-1717

Trionychidae

Aspideretoides

austerus

319a

Kirtland

De-na-zin

incomplete right hypoplastron & other plastron fragments

SMP VP-1844

Trionychidae

Aspideretoides

austerus

319a

Kirtland

De-na-zin

carapace fragments

SMP VP-1868

Trionychidae

Aspideretoides

austerus

382

Kirtland

De-na-zin

costals & nuchal

SMP VP-1959

Trionychidae

Aspideretoides

austerus

210

Kirtland

De-na-zin

large plastron fragment

SMP VP-2376

Trionychidae

Aspideretoides

austerus

389a

Kirtland

De-na-zin

costal & carapace fragments

USNM 6550

Trionychidae

Aspideretoides

austerus

see text

Kirtland

De-na-zin

partial carapace

USNM 12988

Trionychidae

Aspideretoides

austerus

Kirtland

De-na-zin

anterior part of carapace

USNM 12986

Trionychidae

Aspideretoides

ovatus

holotype

Kirtland

Hunter Wash?

partial carapace & plastron

USNM 12987

Trionychidae

Aspideretoides

ovatus

Kirtland

Hunter Wash?

posterior half of carapace

NMMNH P-27779

Trionychidae

Aspideretoides

robustus

4015

Fruitland

Fossil Forest

incomplete right hyoplastron

NMMNH P-27826

Trionychidae

Aspideretoides

robustus

4015

Fruitland

Fossil Forest

incomplete costal & numerous fragments

NMMNH P-27572

Trionychidae

Aspideretoides

robustus

4015

Fruitland

Fossil Forest

fragments

NMMNH P-27628

Trionychidae

Aspideretoides

robustus

4015

Fruitland

Fossil Forest

numerous carapace/plastron fragments

NMMNH P-27633

Trionychidae

Aspideretoides

robustus

4015

Fruitland

Fossil Forest

various carapace/plastron fragments

NMMNH P-27752

Trionychidae

Aspideretoides

robustus

4015

Fruitland

Fossil Forest

incomplete costal & various fragments

NMMNH P-27768

Trionychidae

Aspideretoides

robustus

4015

Fruitland

Fossil Forest

2 plastron fragments

NMMNH P-27777

Trionychidae

Aspideretoides

robustus

4015

Fruitland

Fossil Forest

carapace/plastron fragments

NMMNH P-27941

Trionychidae

Aspideretoides

robustus

1581

Kirtland

Hunter Wash

incomplete costal

NMMNH P-28321

Trionychidae

Aspideretoides

robustus

1882

Fruitland

Fossil Forest

costal fragment

NMMNH P-30066

Trionychidae

Aspideretoides

robustus

4274

Kirtland

Hunter Wash

costal

NMMNH P-30756

Trionychidae

Aspideretoides

robustus

4104

Fruitland/Kirtland

unknown

costal fragments

NMMNH P-39632

Trionychidae

Aspideretoides

robustus

5163

Kirtland

Hunter Wash

2 fused incomplete costals & associated fragments

NMMNH P-49856

UA 11737

Trionychidae

Aspideretoides

robustus

6548

Kirtland?

Hunter Wash?

incomplete costal

NMMNH P-49862

UA 13319

Trionychidae

Aspideretoides

robustus

6560

Kirtland?

Hunter Wash?

incomplete costal

PMU.R 30

Trionychidae

Aspideretoides

robustus

Kirtland

Hunter Wash

nearly complete carapace

PMU.R 31

Trionychidae

Aspideretoides

robustus

Kirtland

Hunter Wash

right hypoplatron & xiphiplastron, plus left hyoplastron, hypoplastron, & xiphiplastron

SMP VP-1612

Trionychidae

Aspideretoides

robustus

400

Kirtland

Hunter Wash

carapace fragments

SMP VP-1667

Trionychidae

Aspideretoides

robustus

398

Kirtland

Hunter Wash

incomplete nuchal, both xiphiplastra, & associated fragments

SMP VP-1679

Trionychidae

Aspideretoides

robustus

400

Kirtland

Hunter Wash

incomplete costal

SMP VP-1681

Trionychidae

Aspideretoides

robustus

400

Kirtland

Hunter Wash

costal fragments

SMP VP-1682

Trionychidae

Aspideretoides

robustus

400

Kirtland

Hunter Wash

nearly complete costal

SMP VP-1685

Trionychidae

Aspideretoides

robustus

396

Kirtland

Hunter Wash

incomplete xiphiplastron

SMP VP-1856

Trionychidae

Aspideretoides

robustus

281

Kirtland

Hunter Wash

carapace fragments

SMP VP-2166

Trionychidae

Aspideretoides

robustus

228

Kirtland

Hunter Wash

carapace fragments

SMP VP-2698

Trionychidae

Aspideretoides

robustus

463

Fruitland

Fossil Forest

carapace fragments, including nuchal & neural

USNM 8350

Trionychidae

Aspideretoides

robustus

Kirtland

Farmington?

left hypoplastron

USNM 8538

Trionychidae

Aspideretoides

robustus

holotype

Kirtland

Hunter Wash

complete carapace

USNM 12989

Trionychidae

Aspideretoides

robustus

Kirtland

Hunter Wash

left hyoplastron & hypoplastron

NMMNH P-33914

Trionychidae

Aspideretoides

cf. A. austerus

4721

Kirtland

De-na-zin

2 incomplete costals

NMMNH P-22751

Trionychidae

Aspideretoides

cf. A. robustus

1675

Fruitland

Fossil Forest

various carapace/plastron fragments

NMMNH P-22934

Trionychidae

Aspideretoides

cf. A. robustus

1790

Kirtland

Hunter Wash

incomplete costal

NMMNH P-22949

Trionychidae

Aspideretoides

cf. A. robustus

1805

Kirtland

Hunter Wash?

2 shell fragments

NMMNH P-27769

Trionychidae

Aspideretoides

cf. A. robustus

4015

Fruitland

Fossil Forest

plastron fragments

NMMNH P-33916

Trionychidae

Aspideretoides

cf. A. robustus

4010

Kirtland

Hunter Wash

nearly complete left xiphiplastron

NMMNH P-28320

Trionychidae

Aspideretoides

cf. A. robustus

1882

Fruitland

Fossil Forest

numerous carapace fragments

NMMNH P-33912

Trionychidae

Aspideretoides

cf. A. robustus

4717

Fruitland

Fossil Forest

numerous carapace/costal fragments

NMMNH P-49848

UA 14397

Trionychidae

Aspideretoides

cf. A. robustus

6547

Kirtland?

Hunter Wash?

shell fragment

NMMNH P-49786

UA 14399

Trionychidae

Aspideretoides

cf. A. robustus

6340

Kirtland?

Hunter Wash?

shell fragment

SMP VP-1525

Trionychidae

Aspideretoides

cf. A. robustus

281

Kirtland

Hunter Wash

costal & other fragments

NMMNH P-19200

Trionychidae

Aspideretoides

sp.

1549

Kirtland

Hunter Wash

incomplete carapace

NMMNH P-22550

Trionychidae

Aspideretoides

sp.

1587

Kirtland

Hunter Wash

carapace fragment

NMMNH P-27829

Trionychidae

Aspideretoides

sp.

4015

Fruitland

Fossil Forest

numerous carapace fragments

NMMNH P-22729

Trionychidae

Aspideretoides

sp.

1665

Kirtland

De-na-zin

incomplete costal

NMMNH P-22735

Trionychidae

Aspideretoides

sp.

1673

Kirtland

Hunter Wash

hypoplastron & associated fragment

NMMNH P-49818

UA 14396

Trionychidae

Aspideretoides

sp.

6559

Kirtland?

Hunter Wash?

2 shell fragments

NMMNH P-49822

UA 11741

Trionychidae

Aspideretoides

sp.

6559

Kirtland?

Hunter Wash?

8 neurals & other fragments

NMMNH P-49832

UA 14412

Trionychidae

Aspideretoides

sp.

6622

Kirtland?

De-na-zin?

costal fragment

SMP VP-1493

Trionychidae

Aspideretoides

sp

319a

Kirtland

De-na-zin

carapace fragments

SMP VP-1519

Trionychidae

Aspideretoides

sp.

365

Kirtland

Hunter Wash

carapace fragments

SMP VP-2045

Trionychidae

Aspideretoides

sp.

388a

Kirtland

De-na-zin

carapace fragments

SMP VP-2601

Trionychidae

Aspideretoides

sp.

319a

Kirtland

De-na-zin

carapace fragment

NMMNH P-49853

UA 14400

Trionychidae

cf. Aspideretoides

sp.

6552

Kirtland?

De-na-zin?

plastron fragment

SMP VP-1368

Trionychidae

cf. Aspideretoides

sp.

349

Kirtland

Hunter Wash

part of carapace

SMP VP-1426

Trionychidae

cf. Aspideretoides

sp.

361

Kirtland

De-na-zin

proximal end of left femur

SMP VP-1594

Trionychidae

cf. Aspideretoides

sp.

398

Kirtland

Hunter Wash

carapace fragments

SMP VP-1595

Trionychidae

cf. Aspideretoides

sp.

398

Kirtland

Hunter Wash

carapace fragments

SMP VP-1810

Trionychidae

cf. Aspideretoides

sp.

409

Kirtland

Hunter Wash

carapace fragment

SMP VP-1949

Trionychidae

cf. Aspideretoides

sp.

228

Kirtland

Hunter Wash

carapace fragment

SMP VP-2803

Trionychidae

cf. Aspideretoides

sp.

450

Fruitland

Fossil Forest

costal, marginal, & other carapace fragments

NMMNH P-22832

Trionychidae

?Aspideretoides

sp.

1714

Kirtland

Hunter Wash

various shell fragments

NMMNH P-26204

Trionychidae

?Aspideretoides

sp.

3228

Kirtland

De-na-zin

costal fragment

NMMNH P-27782

Trionychidae

?Aspideretoides

sp.

4015

Fruitland

Fossil Forest

numerous carapace/plastron fragments

NMMNH P-22627

Trionychidae

Plastomeninae indet.

Plastomeninae indet.

1639

Kirtland

De-na-zin

hypoplastron fragment

NMMNH P-27479

Trionychidae

Plastomeninae indet.

Plastomeninae indet.

4015

Fruitland

Fossil Forest

numerous carapace fragments

SMP VP-742

Trionychidae

Plastomeninae indet.

Plastomeninae indet.

228

Kirtland

Hunter Wash

carapace & plastron fragments

SMP VP-2034

Trionychidae

Plastomeninae indet.

Plastomeninae indet.

421

Kirtland

Hunter Wash

three carapace fragments

SMP VP-2154

Trionychidae

Trionychidae indet.

Trionychidae indet.

281

Kirtland

Hunter Wash

plastron fragment

SMP VP-2156

Trionychidae

Trionychidae indet.

Trionychidae indet.

281

Kirtland

Hunter Wash

carapace fragment

SMP VP-3348

Trionychidae

Trionychidae indet.

Trionychidae indet.

409

Kirtland

Hunter Wash

incomplete costal

NMMNH P-22583

Trionychidae

Trionychidae indet.

Trionychidae indet.

1613

Kirtland

Hunter Wash

carapace fragment

NMMNH P-22804

Trionychidae

Trionychidae indet.

Trionychidae indet.

1699

Kirtland

Hunter Wash

costal fragment & indet. fragment

NMMNH P-26261

Trionychidae

Trionychidae indet.

Trionychidae indet.

3245

Kirtland

De-na-zin

incomplete humerus

NMMNH P-27903

Trionychidae

Trionychidae indet.

Trionychidae indet.

1625

Fruitland

Fossil Forest

carapace & ?dinosaur bone fragment

NMMNH P-28324

Trionychidae

Trionychidae indet.

Trionychidae indet.

1882

Fruitland

Fossil Forest

numerous carapace fragments

NMMNH P-28909

Trionychidae

Trionychidae indet.

Trionychidae indet.

3964

Kirtland

Hunter Wash

incomplete humerus

NMMNH P-32845

Trionychidae

Trionychidae indet.

Trionychidae indet.

4542

Kirtland

De-na-zin

carapace/plastron fragments

NMMNH P-32851

Trionychidae

Trionychidae indet.

Trionychidae indet.

2387

Fruitland

Fossil Forest

carapace/plastron fragments

NMMNH P-32852

Trionychidae

Trionychidae indet.

Trionychidae indet.

2387

Fruitland

Fossil Forest

carapace/plastron fragments

NMMNH P-49825

Trionychidae

Trionychidae indet.

Trionychidae indet.

6558

Kirtland

Hunter Wash

costal fragment

NMMNH P-49855

UA 14399

Trionychidae

Trionychidae indet.

Trionychidae indet.

6552

Kirtland?

De-na-zin?

2 shell fragments

SMP VP-1476

Trionychidae

Trionychidae indet.

Trionychidae indet.

365

Kirtland

Hunter Wash

plastron fragment

SMP VP-1509

Trionychidae

Trionychidae indet.

Trionychidae indet.

228

Kirtland

Hunter Wash

carapace & plastron fragments

SMP VP-1513

Trionychidae

Trionychidae indet.

Trionychidae indet.

365

Kirtland

Hunter Wash

carapace/plastron fragments

SMP VP- SMP VP-1558

Trionychidae

Trionychidae indet.

Trionychidae indet.

389a

Kirtland

De-na-zin

carapace fragment

SMP VP-1567

Trionychidae

Trionychidae indet.

Trionychidae indet.

228

Kirtland

Hunter Wash

carapace fragments

SMP VP-1683

Trionychidae

Trionychidae indet.

Trionychidae indet.

400

Kirtland

Hunter Wash

carapace fragments

SMP VP-1690

Trionychidae

Trionychidae indet.

Trionychidae indet.

409

Kirtland

Hunter Wash

carapace fragment

SMP VP-1691

Trionychidae

Trionychidae indet.

Trionychidae indet.

409

Kirtland

Hunter Wash

carapace fragment

SMP VP-1695

Trionychidae

Trionychidae indet.

Trionychidae indet.

409

Kirtland

Hunter Wash

carapace fragments

SMP VP-1817

Trionychidae

Trionychidae indet.

Trionychidae indet.

365

Kirtland

Hunter Wash

two carapace fragments

SMP VP-1857

Trionychidae

Trionychidae indet.

Trionychidae indet.

365

Kirtland

Hunter Wash

incomplete costal

SMP VP-1917

Trionychidae

Trionychidae indet.

Trionychidae indet.

350

Kirtland

De-na-zin

carapace fragments

SMP VP-1947

Trionychidae

Trionychidae indet.

Trionychidae indet.

228

Kirtland

Hunter Wash

incomplete costal

SMP VP-1996

Trionychidae

Trionychidae indet.

Trionychidae indet.

412

Kirtland

Hunter Wash

costal fragment

SMP VP-2005

Trionychidae

Trionychidae indet.

Trionychidae indet.

350

Kirtland

De-na-zin

four costal fragments

SMP VP-2224

Trionychidae

Trionychidae indet.

Trionychidae indet.

433

Kirtland

Hunter Wash

carapace/plastron fragments

SMP VP-2388

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragments

SMP VP-2403

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace & plastron fragments from multiple individuals

SMP VP-2404

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragments

SMP VP-2423

Trionychidae

Trionychidae indet.

Trionychidae indet.

350

Kirtland

De-na-zin

carapace fragments

SMP VP-2442

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragments from multiple taxa

SMP VP-2460

Trionychidae

Trionychidae indet.

Trionychidae indet.

451

Kirtland

Hunter Wash

carapace fragments

SMP VP-2487

Trionychidae

Trionychidae indet.

Trionychidae indet.

350

Kirtland

De-na-zin

two carapace fragments

SMP VP-2528

Trionychidae

Trionychidae indet.

Trionychidae indet.

452

Kirtland

Hunter Wash

carapace fragments

SMP VP-2530

Trionychidae

Trionychidae indet.

Trionychidae indet.

452

Kirtland

Hunter Wash

neural & carapace fragments

SMP VP-2531

Trionychidae

Trionychidae indet.

Trionychidae indet.

452

Kirtland

Hunter Wash

carapace fragments

SMP VP-2532

Trionychidae

Trionychidae indet.

Trionychidae indet.

452

Kirtland

Hunter Wash

carapace fragments

SMP VP-2541

Trionychidae

Trionychidae indet.

Trionychidae indet.

363a

Kirtland

De-na-zin

carapace fragment

SMP VP-2551

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragment

SMP VP-2586

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragment

SMP VP-2635

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragment

SMP VP-2716

Trionychidae

Trionychidae indet.

Trionychidae indet.

228

Kirtland

Hunter Wash

carapace fragment

SMP VP-2728

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragments

SMP VP-2764

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

incomplete costal

SMP VP-2766

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragments

SMP VP-2772

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

incomplete neural

SMP VP-2819

Trionychidae

Trionychidae indet.

Trionychidae indet.

228

Kirtland

Hunter Wash

carapace neural

SMP VP-2839

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragments

SMP VP-2847

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragments

SMP VP-2852

Trionychidae

Trionychidae indet.

Trionychidae indet.

450

Fruitland

Fossil Forest

carapace fragments & associated indet. bone material

SMP VP-3271

Trionychidae

Trionychidae indet.

Trionychidae indet.

372

Kirtland

Hunter Wash

incomplete neural

SMP VP-3272

Trionychidae

Trionychidae indet.

Trionychidae indet.

372

Kirtland

Hunter Wash

carapace fragment

SMP VP-1135

Cryptodira indet.

Cryptodira indet.

Cryptodira indet.

373

Kirtland

Hunter Wash

incomplete scapula

NMMNH P-22743

Testudines indet.

Testudines indet.

Testudines indet.

1674

Fruitland

Fossil Forest

various shell & bone fragments

NMMNH P-22826

Testudines indet.

Testudines indet.

Testudines indet.

1708

Kirtland

Hunter Wash

various bone fragments

NMMNH P-22921

Testudines indet.

Testudines indet.

Testudines indet.

1782

Kirtland

Hunter Wash

distal end of femur

NMMNH P-22939

Testudines indet.

Testudines indet.

Testudines indet.

1795

Kirtland/Fruitland

unknown

incomplete scapula?

NMMNH P-26246

Testudines indet.

Testudines indet.

Testudines indet.

1708

Kirtland

Hunter Wash

5 pelvic fragments

NMMNH P-26251

Testudines indet.

Testudines indet.

Testudines indet.

3226

Kirtland

De-na-zin

incomplete ?coracoid

NMMNH P-27827

Testudines indet.

Testudines indet.

Testudines indet.

4015

Fruitland

Fossil Forest

incomplete humerus

NMMNH P-22510

Testudines indet.

Testudines indet.

Testudines indet.

1571

Kirtland

Hunter Wash

2 vertebrae

NMMNH P-32823

Testudines indet.

Testudines indet.

Testudines indet.

4010

Kirtland

Hunter Wash

incomplete phalanx

NMMNH P-49847

Testudines indet.

Testudines indet.

Testudines indet.

6546

Kirtland?

Hunter Wash?

shell fragment

NMMNH P-22624

Testudines indet.

Testudines indet.

Testudines indet.

1638

unknown

unknown

pelvis

NMMNH P-37762

Testudines indet.

Testudines indet.

Testudines indet.

5197

Kirtland

De-na-zin

2 carapace fragments

NMMNH P-37787

Testudines indet.

Testudines indet.

Testudines indet.

5197

Kirtland

De-na-zin

14 carapace fragments

NMMNH P-33023

Testudines indet.

Testudines indet.

Testudines indet.

4570

Kirtland

De-na-zin

tibia

SMP VP-817

Testudines indet.

Testudines indet.

Testudines indet.

210

Kirtland

De-na-zin

?posterior part of left ramus

SMP VP-1091

Testudines indet.

Testudines indet.

Testudines indet.

365

Kirtland

Hunter Wash

ungual

SMP VP-1332

Testudines indet.

Testudines indet.

Testudines indet.

385

Kirtland

Hunter Wash

incomplete scapulocoracoid

SMP VP-1420

Testudines indet.

Testudines indet.

Testudines indet.

385

Kirtland

Hunter Wash

girdle & miscellaneous fragments

SMP VP-1471

Testudines indet.

Testudines indet.

Testudines indet.

281

Kirtland

Hunter Wash

pelvic element

SMP VP-1484

Testudines indet.

Testudines indet.

Testudines indet.

281

Kirtland

Hunter Wash

ungual

SMP VP-1530

Testudines indet.

Testudines indet.

Testudines indet.

281

Kirtland

Hunter Wash

ungual & distal end of phalanx

SMP VP-1531

Testudines indet.

Testudines indet.

Testudines indet.

281

Kirtland

Hunter Wash

girdle

SMP VP-1800

Testudines indet.

Testudines indet.

Testudines indet.

228

Kirtland

Hunter Wash

limb fragments

SMP VP-1903

Testudines indet.

Testudines indet.

Testudines indet.

350

Kirtland

De-na-zin

promimal portion of pelvic element (ilium?)

SMP VP-1919

Testudines indet.

Testudines indet.

Testudines indet.

417

Kirtland

Hunter Wash

skull roof?

SMP VP-2000

Testudines indet.

Testudines indet.

Testudines indet.

350

Kirtland

De-na-zin

skull element

SMP VP-2015

Testudines indet.

Testudines indet.

Testudines indet.

420a

Kirtland

De-na-zin

humerus fragment

SMP VP-2035

Testudines indet.

Testudines indet.

Testudines indet.

421

Kirtland

Hunter Wash

unidentified girdle element

SMP VP-2037

Testudines indet.

Testudines indet.

Testudines indet.

421

Kirtland

Hunter Wash

nearly complete left humerus

SMP VP-2084

Testudines indet.

Testudines indet.

Testudines indet.

421

Kirtland

Hunter Wash

incomplete right humerus

SMP VP-2537

Testudines indet.

Testudines indet.

Testudines indet.

316

Kirtland

Hunter Wash

?humerus

SMP VP-2549

Testudines indet.

Testudines indet.

Testudines indet.

450

Fruitland

Fossil Forest

five carapace fragments

SMP VP-2643

Testudines indet.

Testudines indet.

Testudines indet.

450

Fruitland

Fossil Forest

plastron fragment

SMP VP-2681

Testudines indet.

Testudines indet.

Testudines indet.

228

Kirtland

Hunter Wash

incomplete plastron

SMP VP-2739

Testudines indet.

Testudines indet.

Testudines indet.

228

Kirtland

Hunter Wash

incomplete left femur

SMP VP-2747

Testudines indet.

Testudines indet.

Testudines indet.

228

Kirtland

Hunter Wash

cervical vertebra

SMP VP-2768

Testudines indet.

Testudines indet.

Testudines indet.

450

Fruitland

Fossil Forest

distal end of humerus

SMP VP-2777

Testudines indet.

Testudines indet.

Testudines indet.

450

Fruitland

Fossil Forest

?limb osteoderm

SMP VP-2795

Testudines indet.

Testudines indet.

Testudines indet.

450

Fruitland

Fossil Forest

incomplete ?humerus

SMP VP-3343

Testudines indet.

Testudines indet.

Testudines indet.

409

Kirtland

Hunter Wash

incomplete plastron

SMP VP-3344

Testudines indet.

Testudines indet.

Testudines indet.

409

Kirtland

Hunter Wash

incomplete plastron

SMP VP-3373

Testudines indet.

Testudines indet.

Testudines indet.

392

Kirtland

De-na-zin

plastron fragment

SMP VP-3377

Testudines indet.

Testudines indet.

Testudines indet.

461

Kirtland

Hunter Wash

incomplete costal & 2 carapace fragments

SMP VP-3382

Testudines indet.

Testudines indet.

Testudines indet.

450

Fruitland

Fossil Forest

2 plastron fragments

SMP VP-3388

Testudines indet.

Testudines indet.

Testudines indet.

461

Kirtland

Hunter Wash

2 carapace fragments

SMP VP-2381

Testudines indet.

Testudines indet.

Testudines indet.

382

Kirtland

De-na-zin

?girdle element & 2 fragments

SMP VP-2443

Testudines indet.

Testudines indet.

Testudines indet.

421

Kirtland

Hunter Wash

distal & proximal ends of a limb bone

SMP VP-3373

?Testudines indet.

?Testudines indet.

?Testudines indet.

461

Kirtland

Hunter Wash

?carapace/plastron fragments

NMMNH P-28984

various taxa

various taxa

various taxa

4098

Kirtland

unknown

numerous bone fragments

NMMNH P-32887

various taxa

various taxa

various taxa

2387

Fruitland

Fossil Forest

numerous carapace, plastron, axial skeleton fragments

NMMNH P-49857

UA 14835

various taxa

various taxa

various taxa

6552

Kirtland?

De-na-zin?

13 shell fragments

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Sullivan, R.M., Jasinski, S.E., Lucas, S.G. (2013). Re-Assessment of Late Campanian (Kirtlandian) Turtles from the Upper Cretaceous Fruitland and Kirtland Formations, San Juan Basin, New Mexico, USA. In: Brinkman, D., Holroyd, P., Gardner, J. (eds) Morphology and Evolution of Turtles. Vertebrate Paleobiology and Paleoanthropology. Springer, Dordrecht. https://doi.org/10.1007/978-94-007-4309-0_20

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