* Corresponding authors

^a Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences, Tianjin Airport Economic Area, Tianjin, China

^b Key Laboratory Experimental Teratology of the Ministry of Education, Department of Biochemistry and Molecular Biology, School of Basic Medical Sciences, Cheeloo College of Medicine, Shandong University, 44 Wenhua Xi Road, Jinan, Shandong, China
E-mail: sunjinpeng@sdu.edu.cn

^c School of Life Sciences, Tsinghua-Peking Joint Center for Life Sciences, Beijing Advanced Innovation Center for Structural Biology, Tsinghua University, Haidian District, Beijing, China
E-mail: chunlai@tsinghua.edu.cn

^d Department of Physiology and Pathophysiology, School of Basic Medical Sciences, Peking University, Key Laboratory of Molecular Cardiovascular Science, Ministry of Education, Haidian District, Beijing, China

^e Institute of Biophysics, Chinese Academy of Sciences, Chaoyang District, Beijing, China

^f University of the Chinese Academy of Sciences (UCAS), Shijingshan District, Beijing, China

^g Shenzhen Institute of Transfusion Medicine, Shenzhen Blood Center, Futian District, Shenzhen, China

^h School of Life Sciences, University of Science and Technology of China, Baohe District, Anhui, China

ⁱ School of Biological Science and Technology, University of Jinan, Jinan, Shandong, China