* Corresponding authors

^a School of Engineering and Applied Sciences, Harvard University, Cambridge, MA 02138, USA

^b School of Mechatronics Engineering, Harbin Institute of Technology, Harbin 150001, China

^c Department of Cell Biology, Key Laboratory of Cell Biology, Ministry of Public Health, and Key Laboratory of Medical Cell Biology, Ministry of Education, China Medical University, Shenyang 110001, China

^d Chemical and Biological Engineering Department, Montana State University, Bozeman, MT 59717, USA

^e Broad Institute of MIT and Harvard 7 Cambridge Center, MA 02142, USA

^f Department of Microbiology and Immunology, University of Michigan, Ann Arbor, MI 48109, USA

^g Applied Physics Laboratory, Johns Hopkins University, Laurel, Maryland 11100, USA

^h Department of Biological Sciences, University of Pittsburgh, Pittsburgh, PA 15260, USA

ⁱ Department of Physics, Harvard University, Cambridge, MA 02138, USA
E-mail: weitz@seas.harvard.edu